Pellaea wrightiana |
Pellaea breweri |
|
---|---|---|
Wright's cliffbrake |
Brewer's cliff-brake, Brewer's cliffbrake fern |
|
Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
compact, ascending, stout, 5–10 mm diam.; scales uniformly reddish brown, linear-subulate, 0.1–0.3 mm wide, thin, margins sinuous, nearly entire. |
Leaves | monomorphic, clustered on stem, 6–40 cm; croziers sparsely villous. |
monomorphic, clustered on stem, 2.5–20 cm; croziers sparsely villous. |
Petiole | dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. |
brown, lustrous, rounded adaxially, with prominent articulation lines near base. |
Blade | linear-oblong, 2-pinnate proximally, 1.5–5 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. |
linear-oblong, pinnate-pinnatifid proximally, 1–4 cm wide; rachis brown proximally, green distally, straight, rounded adaxially, glabrous to sparsely villous. |
Ultimate segments | narrowly oblong, 5–20 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate; apex mucronate. |
lanceolate-deltate, 5–25 mm, herbaceous, glabrous; margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, erose-denticulate; apex obtuse or rounded. |
Pinnae | perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–9 ultimate segments; costae straight, 2–20 mm, usually shorter than ultimate segments. |
ascending or perpendicular to rachis, decurrent on rachis, deeply 2-lobed (mitten-shaped) near base of leaf; costae absent. |
Veins | of ultimate segments obscure. |
of ultimate segments evident. |
Sporangia | long-stalked, containing 64 spores, intermixed with sparse farina-producing glands. |
sessile or subsessile, containing 64 spores, not intermixed with farina-producing glands. |
2n | = 116. |
= 58. |
Pellaea wrightiana |
Pellaea breweri |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates | Cliffs and rocky slopes, on a variety of substrates including granite and limestone |
Elevation | 300–2900 m (1000–9500 ft) | 1600–3800 m (5200–12500 ft) |
Distribution |
AZ; CO; NC; NM; OK; TX; UT; n Mexico
|
CA; CO; ID; MT; NV; OR; UT; WA; WY
|
Discussion | W. H. Wagner Jr. (1965) suggested that Pellaea wrightiana was a fertile allotetraploid hybrid between P. truncata (as P. longimucronata) and P. ternifolia. This hypothesis has been confirmed by isozyme analyses (M. D. Windham 1988). Pellaea wrightiana is therefore treated as a distinct species rather than a variety of P. ternifolia. This tetraploid species hybridizes with P. truncata and P. ternifolia subsp. arizonica to produce sterile triploids and tetraploids with intermediate morphology and malformed spores. Pellaea wrightiana has also hybridized with P. atropurpurea to form a rare apogamous pentaploid known only from western Oklahoma. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea breweri is distinguished from other North American taxa (except for some populations of P. glabella) by the presence of prominent articulation lines near the base of the petiole. The leaves are easily detached, and many herbarium specimens consist of separate leaves and stems, the latter covered with petiole bases of approximately equal length. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea |
Sibling taxa | ||
Synonyms | P. ternifolia var. wrightiana | |
Name authority | Hooker: Sp. Fil. 2: 142. (1858) | D. C. Eaton: Proc. Amer. Acad. Arts 6: 555. (1865) |
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