Pellaea truncata |
Pellaea bridgesii |
|
---|---|---|
cliff brake, spiny cliff-brake |
Bridges' cliff-brake |
|
Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
compact, ascending, stout, 5–10 mm diam.; scales mostly weakly bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown, thin, margins lighter, thin, denticulate to entire. |
Leaves | somewhat dimorphic, sterile leaves shorter and less divided than fertile leaves, clustered on stems, 8–40 cm; croziers sparsely villous. |
monomorphic, clustered on stem, 7–30 cm; croziers nearly glabrous. |
Petiole | dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. |
dark brown, lustrous, rounded adaxially, without prominent articulation lines. |
Blade | ovate-deltate, usually 2-pinnate proximally, 4–18 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. |
linear, 1-pinnate, 1.5–4 cm wide; rachis brown throughout, straight, rounded adaxially, glabrous. |
Ultimate segments | narrowly oblong, 4–10 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, nearly entire; apex mucronate. |
broadly ovate to elliptic, 7–20 mm, leathery, glabrous; margins plane, not recurved, not covering abaxial surface, borders whitish, entire; apex obtuse to rounded. |
Pinnae | perpendicular to rachis to slightly ascending, not decurrent on rachis, usually with 9–25 ultimate segments; costae straight, 20–70 mm, much longer than fertile ultimate segments. |
perpendicular to slightly ascending, usually not decurrent on rachis, simple and unlobed; costae absent. |
Veins | of ultimate segments obscure. |
of ultimate segments obscure. |
Sporangia | long-stalked, containing 64 spores, intermixed with abundant farina-producing glands. |
sessile or subsessile, containing 64 spores, intermixed with abundant farina-producing glands. |
2n | = 58. |
= 58. |
Pellaea truncata |
Pellaea bridgesii |
|
Phenology | Sporulating late spring–fall. | Sporulating summer–fall. |
Habitat | Cliffs and rocky slopes, on various substrates but rarely observed on limestone | Rocky slopes and cliffs, on granitic substrates |
Elevation | 600–2500 m (2000–8200 ft) | 1200–3600 m (3900–11800 ft) |
Distribution |
AZ; CA; CO; NM; NV; TX; UT; n Mexico
|
CA; ID; NV; OR
|
Discussion | Most manuals refer to Pellaea truncata as P. longimucronata, a name shown to be invalid by A. Cronquist et al. (1972+, vol. 1). Populations located near the range of P. mucronata in the Mojave Desert are often difficult to identify because of the subtlety of the characters involved and an apparent tendency to produce sterile (and possibly fertile) hybrids. Morphologically intermediate hybrids between P. truncata and P. wrightiana are common in regions where the ranges of the two species overlap, but these are easily identified by their malformed spores. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The morphology of Pellaea bridgesii is so distinctive that its sectional (and even generic) placement in Pellaea has long been a source of contention. W. H. Wagner Jr. et al. (1983) documented the existence of sterile diploid hybrids (called P. × glaciogena) between P. bridgesii and P. mucronata (see reticulogram), suggesting that P. bridgesii is most closely related to members of sect. Pellaea. In addition to the more obvious characters mentioned above, P. bridgesii is distinguished from other North American species (except P. ternifolia) by its anastomosing veins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea |
Sibling taxa | ||
Synonyms | P. longimucronata, P. wrightiana var. longimucronata | |
Name authority | Goodding: Muhlenbergia 8: 94. (1912) | Hooker: Sp. Fil. 2: 238, plate 142b. (1858) |
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