Pellaea ternifolia |
Pellaea breweri |
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trans-Pecos cliffbrake |
Brewer's cliff-brake, Brewer's cliffbrake fern |
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Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
compact, ascending, stout, 5–10 mm diam.; scales uniformly reddish brown, linear-subulate, 0.1–0.3 mm wide, thin, margins sinuous, nearly entire. |
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Leaves | monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. |
monomorphic, clustered on stem, 2.5–20 cm; croziers sparsely villous. |
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Petiole | black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
brown, lustrous, rounded adaxially, with prominent articulation lines near base. |
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Blade | linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. |
linear-oblong, pinnate-pinnatifid proximally, 1–4 cm wide; rachis brown proximally, green distally, straight, rounded adaxially, glabrous to sparsely villous. |
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Ultimate segments | linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. |
lanceolate-deltate, 5–25 mm, herbaceous, glabrous; margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, erose-denticulate; apex obtuse or rounded. |
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Pinnae | perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. |
ascending or perpendicular to rachis, decurrent on rachis, deeply 2-lobed (mitten-shaped) near base of leaf; costae absent. |
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Veins | of ultimate segments obscure. |
of ultimate segments evident. |
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Sporangia | long-stalked, containing 64 spores, not intermixed with farina-producing glands. |
sessile or subsessile, containing 64 spores, not intermixed with farina-producing glands. |
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2n | = 58. |
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Pellaea ternifolia |
Pellaea breweri |
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Phenology | Sporulating summer–fall. | |||||||||
Habitat | Cliffs and rocky slopes, on a variety of substrates including granite and limestone | |||||||||
Elevation | 1600–3800 m (5200–12500 ft) | |||||||||
Distribution |
AZ; TX; Mexico; Central America; South America; Pacific Islands in Hawaii
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CA; CO; ID; MT; NV; OR; UT; WA; WY
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Discussion | Subspecies 3 (3 in the flora). Pellaea ternifolia is represented in the flora by three morphologically and chromosomally distinct taxa. These discrete genetic entities also show a tendency toward geographic isolation and are treated here as subspecies. Diploid populations referred to P. ternifolia subsp. ternifolia are scattered from Texas through Mexico to South America. The pubescent tetraploid (P. ternifolia subsp. villosa) follows the Sierra Madre Oriental from Puebla, Mexico, north to Texas; the glabrous tetraploid (P. ternifolia subsp. arizonica) occurs in Arizona, Texas, and northern Mexico. Isozyme and chromosome studies suggest that both tetraploids are segmental allopolyploids produced by hybridization between subsp. ternifolia and other (as yet unidentified) diploid elements within P. ternifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea breweri is distinguished from other North American taxa (except for some populations of P. glabella) by the presence of prominent articulation lines near the base of the petiole. The leaves are easily detached, and many herbarium specimens consist of separate leaves and stems, the latter covered with petiole bases of approximately equal length. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2, p. 180. | FNA vol. 2. | ||||||||
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Pteris ternifolia | |||||||||
Name authority | (Cavanilles) Link: Fil. Spec. 59. (1841) | D. C. Eaton: Proc. Amer. Acad. Arts 6: 555. (1865) | ||||||||
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