Pellaea ternifolia |
Pellaea brachyptera |
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trans-Pecos cliffbrake |
Sierra cliff-brake, Sierran cliffbrake |
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Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown to black, thick, margins brown, thin, dentate. |
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Leaves | monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. |
monomorphic, clustered on stem, 8–40 cm; croziers sparsely villous. |
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Petiole | black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. |
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Blade | linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. |
linear-oblong, 2-pinnate proximally, 1–4 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. |
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Ultimate segments | linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. |
linear, 5–20 mm, leathery, glabrous; margins on fertile segments strongly revolute, covering more than 1/2 abaxial surface, borders greenish, crenate; apex mucronate. |
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Pinnae | perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. |
strongly ascending, not decurrent on rachis, usually with 5–11 ultimate segments; costae straight, 5–20 mm, usually shorter than ultimate segments. |
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Veins | of ultimate segments obscure. |
of ultimate segments obscure. |
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Sporangia | long-stalked, containing 64 spores, not intermixed with farina-producing glands. |
short-stalked, containing 64 spores, intermixed with abundant farina-producing glands. |
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Pellaea ternifolia |
Pellaea brachyptera |
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Phenology | Sporulating summer–fall. | |||||||||
Habitat | Cliffs and rocky slopes, usually on igneous substrates, occasionally on serpentine | |||||||||
Elevation | 900–2700 m (3000–8900 ft) | |||||||||
Distribution |
AZ; TX; Mexico; Central America; South America; Pacific Islands in Hawaii
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CA; OR; WA
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Discussion | Subspecies 3 (3 in the flora). Pellaea ternifolia is represented in the flora by three morphologically and chromosomally distinct taxa. These discrete genetic entities also show a tendency toward geographic isolation and are treated here as subspecies. Diploid populations referred to P. ternifolia subsp. ternifolia are scattered from Texas through Mexico to South America. The pubescent tetraploid (P. ternifolia subsp. villosa) follows the Sierra Madre Oriental from Puebla, Mexico, north to Texas; the glabrous tetraploid (P. ternifolia subsp. arizonica) occurs in Arizona, Texas, and northern Mexico. Isozyme and chromosome studies suggest that both tetraploids are segmental allopolyploids produced by hybridization between subsp. ternifolia and other (as yet unidentified) diploid elements within P. ternifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The distinctive Pellaea brachyptera reportedly hybridizes with P. mucronata (A. F. Tryon 1957; D. B. Lellinger 1985); the hybrids are morphologically intermediate plants with malformed spores. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2, p. 180. | FNA vol. 2. | ||||||||
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Pteris ternifolia | Platyloma brachyptera | ||||||||
Name authority | (Cavanilles) Link: Fil. Spec. 59. (1841) | (T. Moore) Baker: in Hooker & Baker, Syn. Fil. ed. 2 477. (1874) | ||||||||
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