Pellaea ternifolia |
Pellaea andromedifolia |
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trans-Pecos cliffbrake |
coffee cliffbrake, coffee fern |
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Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
creeping, horizontal, slender, 2–4 mm diam.; scales mostly bicolored, narrowly lanceolate, largest scales 0.3–0.8 mm wide, centers black, thick, margins brown, thin, irregularly dentate. |
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Leaves | monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. |
monomorphic, scattered along stem, 10–60 cm; croziers not conspicuously pubescent, densely scaly. |
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Petiole | black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
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Blade | linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. |
elongate-deltate, usually 3-pinnate proximally, 3–20 cm wide; rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, glabrous or pubescent. |
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Ultimate segments | linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. |
elliptic to ovate, 3–15 mm, somewhat herbaceous, glabrous to sparsely pubescent abaxially; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, entire; apex retuse to rounded. |
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Pinnae | perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. |
ascending or perpendicular to rachis, not decurrent on rachis, with 8–50 ultimate segments; costae usually straight, 15–140 mm, longer than ultimate segments. |
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Veins | of ultimate segments obscure. |
of ultimate segments evident. |
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Sporangia | long-stalked, containing 64 spores, not intermixed with farina-producing glands. |
short-stalked, containing 64 or 32 spores, not intermixed with farina-producing glands. |
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2n | = 58; n = 2n = 87, 116, apogamous. |
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Pellaea ternifolia |
Pellaea andromedifolia |
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Phenology | Sporulating late spring–summer. | |||||||||
Habitat | Rocky slopes and ledges, usually on igneous substrates | |||||||||
Elevation | 0–1500 m (0–4900 ft) | |||||||||
Distribution |
AZ; TX; Mexico; Central America; South America; Pacific Islands in Hawaii
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CA; OR; Mexico in Baja California
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Discussion | Subspecies 3 (3 in the flora). Pellaea ternifolia is represented in the flora by three morphologically and chromosomally distinct taxa. These discrete genetic entities also show a tendency toward geographic isolation and are treated here as subspecies. Diploid populations referred to P. ternifolia subsp. ternifolia are scattered from Texas through Mexico to South America. The pubescent tetraploid (P. ternifolia subsp. villosa) follows the Sierra Madre Oriental from Puebla, Mexico, north to Texas; the glabrous tetraploid (P. ternifolia subsp. arizonica) occurs in Arizona, Texas, and northern Mexico. Isozyme and chromosome studies suggest that both tetraploids are segmental allopolyploids produced by hybridization between subsp. ternifolia and other (as yet unidentified) diploid elements within P. ternifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea andromedifolia comprises three cytotypes: a sexually reproducing diploid, an apogamous triploid, and an apogamous tetraploid. Isozyme studies by G. J. Gastony and L. D. Gottlieb (1985) suggested that the apogamous triploid is an autopolyploid derived from sexual diploid populations. The apogamous tetraploid apparently resulted from hybridization between diploid and triploid individuals. These cytotypes have not been formally recognized as subspecies because their ranges seem to overlap extensively and because the ploidy level of the type collection of P. andromedifolia is not known. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2, p. 180. | FNA vol. 2. | ||||||||
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Pteris ternifolia | Pteris andromedifolia, P. andromedifolia var. pubescens | ||||||||
Name authority | (Cavanilles) Link: Fil. Spec. 59. (1841) | (Kaulfuss) Fée | ||||||||
Web links |