Pellaea ternifolia |
Pellaea |
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trans-Pecos cliffbrake |
cliff-brake |
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Habit | Plants usually on rock. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. |
compact to long-creeping, ascending to horizontal, usually branched; scales brown to tan or often bicolored with dark, central stripe and lighter margins, linear-subulate to lanceolate (rarely ovate), margins dentate, erose, or entire. |
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Leaves | monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. |
monomorphic to somewhat dimorphic, clustered to widely scattered, 2–100 cm. |
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Petiole | black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
brown, black, straw-colored, or gray, rounded, flattened or with single longitudinal groove adaxially, glabrous or pubescent, usually with a few scales at base, with single vascular bundle. |
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Blade | linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. |
linear to ovate-deltate, 1–4-pinnate proximally, leathery or rarely somewhat herbaceous, abaxially glabrous, pubescent, or with hairlike scales scattered along costae, adaxially usually glabrous, dull, not striate; rachis straight or flexuous. |
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Ultimate segments | linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. |
of blade usually stalked and free from costae, elliptic, lanceolate to linear, usually more than 4 mm wide; base rounded, truncate, or cordate; stalks often lustrous and dark colored; segment margins reflexed to form confluent, poorly defined, false indusia extending entire length of segment. |
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Pinnae | perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. |
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Veins | of ultimate segments obscure. |
of ultimate segments free or rarely anastomosing, usually obscure, pinnately branched and divergent distally. |
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False indusia | greenish to whitish, narrow, clearly marginal, often concealing the sporangia. |
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Sporangia | long-stalked, containing 64 spores, not intermixed with farina-producing glands. |
scattered along veins near segment margins, containing 32 or 64 spores, often intermixed with glands, farina-producing. |
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Spores | brown to tan (rarely yellow), tetrahedral-globose, rugose or cristate, lacking prominent equatorial ridge. |
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x | = 29. |
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Pellaea ternifolia |
Pellaea |
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Distribution |
AZ; TX; Mexico; Central America; South America; Pacific Islands in Hawaii
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Most in the Western Hemisphere; a small number in Asia; Africa; the Pacific Islands; and Australia |
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Discussion | Subspecies 3 (3 in the flora). Pellaea ternifolia is represented in the flora by three morphologically and chromosomally distinct taxa. These discrete genetic entities also show a tendency toward geographic isolation and are treated here as subspecies. Diploid populations referred to P. ternifolia subsp. ternifolia are scattered from Texas through Mexico to South America. The pubescent tetraploid (P. ternifolia subsp. villosa) follows the Sierra Madre Oriental from Puebla, Mexico, north to Texas; the glabrous tetraploid (P. ternifolia subsp. arizonica) occurs in Arizona, Texas, and northern Mexico. Isozyme and chromosome studies suggest that both tetraploids are segmental allopolyploids produced by hybridization between subsp. ternifolia and other (as yet unidentified) diploid elements within P. ternifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea in the broad sense is a diverse, poorly defined assemblage of xeric-adapted ferns (A. R. Smith 1981). Relationships among the North American, neotropical, and Eastern Hemisphere species are unclear, and it seems likely that the genus, as broadly construed by E. B. Copeland (1947) and R. M. Tryon and A. F. Tryon (1982), is polyphyletic. The species included here in Pellaea belong to a closely knit alliance that is usually recognized as a distinct section (sect. Pellaea). Although the inclusion of P. bridgesii in this group has been questioned (A. F. Tryon 1957), W. H. Wagner Jr. et al. (1983) have shown that the aberrant morphology of this species is simply an extreme expression of evolutionary trends commonly encountered in sect. Pellaea. Among Western Hemisphere cheilanthoid ferns, species of Pellaea show clear morphologic, chromosomal, and biochemical affinities to Argyrochosma and members of the Cheilanthes alabamensis complex. In fact, the glabrous species of Argyrochosma (A. jonesii and A. microphylla) are commonly misidentified as Pellaea. These species are easily recognizable, however, because they have a combination of concolored stem scales and small ultimate segments (less than 4 mm wide). Species ca. 40 (15 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2, p. 180. | FNA vol. 2. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Pteridaceae > Pellaea | Pteridaceae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Pteris ternifolia | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Cavanilles) Link: Fil. Spec. 59. (1841) | Link: Fil. Spec. 59. (1841) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |
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