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creeping cliff brake, intermediate cliffbrake

lyngholm's cliffbrake

Stems

creeping, horizontal, slender, 2–4 mm diam.;

scales mostly bicolored, narrowly lanceolate, largest scales 0.3–0.8 mm wide, centers black, thick, margins brown, thin, irregularly dentate.

compact, ascending, stout, 5–10 mm diam.;

scales uniformly brown or tan, linear-subulate, 0.1–0.3 mm wide, thin, margins entire to denticulate.

Leaves

monomorphic, widely scattered along stem, 12–50 cm;

croziers pubescent and bearing a few scales.

somewhat dimorphic, sterile leaves shorter and less divided than fertile leaves, clustered on stem, 10–30 cm;

croziers villous.

Petiole

straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.

dark brown to reddish purple, lustrous, rounded adaxially, without prominent articulation lines.

Blade

ovate to elongate-deltate, usually 2-pinnate proximally, 4–20 cm wide;

rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, ± pubescent.

elongate-deltate to ovate, 2-pinnate proximally, 5–15 cm wide;

rachis brown or reddish purple throughout, straight, often slightly flattened adaxially, sparsely villous with long, divergent hairs.

Ultimate segments

ovate to elliptic, 5–15 mm, leathery, glabrous or usually puberulent abaxially;

margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, nearly entire;

apex obtuse to slightly mucronate.

oblong-lanceolate, 7–25 mm, leathery, sparsely villous abaxially near midrib;

margins usually recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, crenulate;

apex slightly mucronate.

Pinnae

perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 7–21 ultimate segments;

costae straight to slightly flexuous, 20–100 mm, longer than ultimate segments.

perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 7–15 ultimate segments;

costae straight, 25–80 mm, usually longer than ultimate segments.

Veins

of ultimate segments obscure.

of ultimate segments obscure.

Sporangia

short-stalked, containing 32 spores, not intermixed with farina-producing glands.

long-stalked, containing 32 spores, not intermixed with farina-producing glands.

n

= 2n = 87, 116, apogamous.

Pellaea intermedia

Pellaea lyngholmii

Phenology Sporulating summer–fall. Sporulating summer–fall.
Habitat Rocky slopes and ledges, on a variety of substrates, including limestone and granite Rocky slopes and ledges, usually on sandstone
Elevation 300–2400 m (1000–7900 ft) 1200–1800 m (3900–5900 ft)
Distribution
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Plants of Pellaea intermedia in the flora are apogamous triploids and tetraploids; a sexual diploid cytotype has been found near Saltillo, Mexico (A. F. Tryon 1968). Given the high degree of morphologic similarity among the three cytotypes, the North American polyploids probably were derived from the Mexican diploid through autopolyploidy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pellaea lyngholmii is an apogamous tetraploid that arose through hybridization between P. atropurpurea and P. truncata (M. D. Windham 1993). It is most often confused with P. atropurpurea, from which P. lyngholmii differs in having sparsely villous rachises, smaller and more numerous ultimate segments, and spores usually more than 62 µm in diameter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Pellaea Pteridaceae > Pellaea
Sibling taxa
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. intermedia, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
Synonyms P. intermedia var. pubescens
Name authority Mettenius ex Kuhn: Linnaea 38: 84. (1869) Windham: Contr. Univ. Michigan Herb. 19: 40. (1993)
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