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creeping cliff brake, intermediate cliffbrake

Stems

creeping, horizontal, slender, 2–4 mm diam.;

scales mostly bicolored, narrowly lanceolate, largest scales 0.3–0.8 mm wide, centers black, thick, margins brown, thin, irregularly dentate.

Leaves

monomorphic, widely scattered along stem, 12–50 cm;

croziers pubescent and bearing a few scales.

Petiole

straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.

Blade

ovate to elongate-deltate, usually 2-pinnate proximally, 4–20 cm wide;

rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, ± pubescent.

Ultimate segments

ovate to elliptic, 5–15 mm, leathery, glabrous or usually puberulent abaxially;

margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, nearly entire;

apex obtuse to slightly mucronate.

Pinnae

perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 7–21 ultimate segments;

costae straight to slightly flexuous, 20–100 mm, longer than ultimate segments.

Veins

of ultimate segments obscure.

Sporangia

short-stalked, containing 32 spores, not intermixed with farina-producing glands.

n

= 2n = 87, 116, apogamous.

Pellaea intermedia

Phenology Sporulating summer–fall.
Habitat Rocky slopes and ledges, on a variety of substrates, including limestone and granite
Elevation 300–2400 m [1000–7900 ft]
Discussion

Plants of Pellaea intermedia in the flora are apogamous triploids and tetraploids; a sexual diploid cytotype has been found near Saltillo, Mexico (A. F. Tryon 1968). Given the high degree of morphologic similarity among the three cytotypes, the North American polyploids probably were derived from the Mexican diploid through autopolyploidy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Distribution
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Parent taxa Pteridaceae > Pellaea
Sibling taxa
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
Synonyms P. intermedia var. pubescens
Name authority Mettenius ex Kuhn: Linnaea 38: 84. (1869)
Source Flora of North America vol. 2.
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