Pellaea bridgesii |
Pellaea glabella |
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Bridges' cliff-brake |
pelléade glabre, simple cliff-brake, slender cliff-brake, smooth cliff-brake, smooth cliffbrake fern |
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Stems | compact, ascending, stout, 5–10 mm diam.; scales mostly weakly bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown, thin, margins lighter, thin, denticulate to entire. |
compact, ascending, stout, 5–10 mm diam.; scales uniformly reddish brown, linear-subulate, 0.1–0.3 mm wide, thin, margins sinuous, entire to denticulate. |
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Leaves | monomorphic, clustered on stem, 7–30 cm; croziers nearly glabrous. |
monomorphic, clustered on stem, 2–40 cm; croziers sparsely villous. |
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Petiole | dark brown, lustrous, rounded adaxially, without prominent articulation lines. |
brown, lustrous, rounded adaxially, occasionally with prominent articulation lines near base. |
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Blade | linear, 1-pinnate, 1.5–4 cm wide; rachis brown throughout, straight, rounded adaxially, glabrous. |
linear-oblong to ovate-lanceolate, 1–2-pinnate proximally, 1–8 cm wide; rachis brown throughout, straight, rounded adaxially, nearly glabrous. |
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Ultimate segments | broadly ovate to elliptic, 7–20 mm, leathery, glabrous; margins plane, not recurved, not covering abaxial surface, borders whitish, entire; apex obtuse to rounded. |
oblong-lanceolate, 5–20 mm, leathery to herbaceous, glabrous except for occasional hairlike scales abaxially near midrib; margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, erose-denticulate; apex obtuse. |
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Pinnae | perpendicular to slightly ascending, usually not decurrent on rachis, simple and unlobed; costae absent. |
somewhat ascending, decurrent on rachis, usually with 3–7 lobes or ultimate segments; costae when present straight, 1–50 mm, often shorter than ultimate segments. |
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Veins | of ultimate segments obscure. |
of ultimate segments usually obscure. |
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Sporangia | sessile or subsessile, containing 64 spores, intermixed with abundant farina-producing glands. |
long-stalked, containing 32 or 64 spores, not intermixed with farina-producing glands. |
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2n | = 58. |
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Pellaea bridgesii |
Pellaea glabella |
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Phenology | Sporulating summer–fall. | |||||||||||||
Habitat | Rocky slopes and cliffs, on granitic substrates | |||||||||||||
Elevation | 1200–3600 m (3900–11800 ft) | |||||||||||||
Distribution |
CA; ID; NV; OR
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AR; AZ; CO; CT; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MT; ND; NE; NM; NY; OH; OK; PA; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; only in the flora
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Discussion | The morphology of Pellaea bridgesii is so distinctive that its sectional (and even generic) placement in Pellaea has long been a source of contention. W. H. Wagner Jr. et al. (1983) documented the existence of sterile diploid hybrids (called P. × glaciogena) between P. bridgesii and P. mucronata (see reticulogram), suggesting that P. bridgesii is most closely related to members of sect. Pellaea. In addition to the more obvious characters mentioned above, P. bridgesii is distinguished from other North American species (except P. ternifolia) by its anastomosing veins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea glabella includes four geographically and genetically isolated taxa treated here as subspecies. D. B. Lellinger (1985) recognized three species in this difficult group, but isozyme analyses (G. J. Gastony 1988) showed that one of these (P. suksdorfiana) is an autotetraploid derivative of the diploid known as P. occidentalis. As a result, Gastony recognized just two species: P. glabella (with two varieties) and P. occidentalis (with two subspecies). The few morphologic features that distinguish these taxa, however, are subtle and environmentally plastic, and the isozyme data indicate that they are less divergent genetically than any other pair of Pellaea species in North America. Therefore, a more conservative taxonomic treatment seems warranted. Subspecies 4. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||||||||||
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Name authority | Hooker: Sp. Fil. 2: 238, plate 142b. (1858) | Mettenius ex Kuhn: Linnaea 36: 87. (1869) | ||||||||||||
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