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Bridges' cliff-brake

heartleaf cliffbrake

Stems

compact, ascending, stout, 5–10 mm diam.;

scales mostly weakly bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown, thin, margins lighter, thin, denticulate to entire.

compact, ascending, stout, 6–10 mm diam.;

scales uniformly orange-brown and thin, lanceolate to ovate, largest scales 0.3–1 mm wide, margins dentate.

Leaves

monomorphic, clustered on stem, 7–30 cm;

croziers nearly glabrous.

somewhat dimorphic, sterile leaves shorter than fertile leaves, clustered on stem, 15–50 cm;

croziers not conspicuously pubescent, densely scaly.

Petiole

dark brown, lustrous, rounded adaxially, without prominent articulation lines.

straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.

Blade

linear, 1-pinnate, 1.5–4 cm wide;

rachis brown throughout, straight, rounded adaxially, glabrous.

ovate-deltate, 2-pinnate proximally, 5–20 cm wide;

rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, glabrous.

Ultimate segments

broadly ovate to elliptic, 7–20 mm, leathery, glabrous;

margins plane, not recurved, not covering abaxial surface, borders whitish, entire;

apex obtuse to rounded.

round-cordate to deltate-cordate, 5–15 mm, herbaceous to leathery, glabrous or puberulent;

margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, crenulate;

apex rounded or retuse.

Pinnae

perpendicular to slightly ascending, usually not decurrent on rachis, simple and unlobed;

costae absent.

perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–15 ultimate segments;

costae straight to slightly flexuous, 25–100 mm, longer than ultimate segments.

Veins

of ultimate segments obscure.

of ultimate segments usually evident.

Sporangia

sessile or subsessile, containing 64 spores, intermixed with abundant farina-producing glands.

short-stalked, containing 64 spores, not intermixed with farina-producing glands.

2n

= 58.

= 58.

Pellaea bridgesii

Pellaea cordifolia

Phenology Sporulating summer–fall. Sporulating summer–fall.
Habitat Rocky slopes and cliffs, on granitic substrates Rocky slopes and ledges, usually on volcanic substrates
Elevation 1200–3600 m (3900–11800 ft) 1000–2500 m (3300–8200 ft)
Distribution
from FNA
CA; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico
[BONAP county map]
Discussion

The morphology of Pellaea bridgesii is so distinctive that its sectional (and even generic) placement in Pellaea has long been a source of contention. W. H. Wagner Jr. et al. (1983) documented the existence of sterile diploid hybrids (called P. × glaciogena) between P. bridgesii and P. mucronata (see reticulogram), suggesting that P. bridgesii is most closely related to members of sect. Pellaea. In addition to the more obvious characters mentioned above, P. bridgesii is distinguished from other North American species (except P. ternifolia) by its anastomosing veins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The diploid Pellaea cordifolia has often been treated as a variety of the Central American and South American apogamous triploid, P. sagittata. The two taxa are distinguished by a number of qualitative morphologic features (A. R. Smith 1980), and it seems unlikely that they represent cytotypes of a single species. A. F. Tryon (1957) suggested that P. sagittata may have originated through hybridization between P. ovata and P. cordifolia (as P. sagittata var. cordata).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Pellaea Pteridaceae > Pellaea
Sibling taxa
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. cordifolia, P. gastonyi, P. glabella, P. intermedia, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. gastonyi, P. glabella, P. intermedia, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
Synonyms Adiantum cordifolium, P. cardiomorpha, P. sagittata var. cordata
Name authority Hooker: Sp. Fil. 2: 238, plate 142b. (1858) (Sessé & Mociño) A. R. Smith: Amer. Fern J. 70: 26. (1980)
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