Pellaea bridgesii |
Pellaea cordifolia |
|
---|---|---|
Bridges' cliff-brake |
heartleaf cliffbrake |
|
Stems | compact, ascending, stout, 5–10 mm diam.; scales mostly weakly bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown, thin, margins lighter, thin, denticulate to entire. |
compact, ascending, stout, 6–10 mm diam.; scales uniformly orange-brown and thin, lanceolate to ovate, largest scales 0.3–1 mm wide, margins dentate. |
Leaves | monomorphic, clustered on stem, 7–30 cm; croziers nearly glabrous. |
somewhat dimorphic, sterile leaves shorter than fertile leaves, clustered on stem, 15–50 cm; croziers not conspicuously pubescent, densely scaly. |
Petiole | dark brown, lustrous, rounded adaxially, without prominent articulation lines. |
straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. |
Blade | linear, 1-pinnate, 1.5–4 cm wide; rachis brown throughout, straight, rounded adaxially, glabrous. |
ovate-deltate, 2-pinnate proximally, 5–20 cm wide; rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, glabrous. |
Ultimate segments | broadly ovate to elliptic, 7–20 mm, leathery, glabrous; margins plane, not recurved, not covering abaxial surface, borders whitish, entire; apex obtuse to rounded. |
round-cordate to deltate-cordate, 5–15 mm, herbaceous to leathery, glabrous or puberulent; margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, crenulate; apex rounded or retuse. |
Pinnae | perpendicular to slightly ascending, usually not decurrent on rachis, simple and unlobed; costae absent. |
perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–15 ultimate segments; costae straight to slightly flexuous, 25–100 mm, longer than ultimate segments. |
Veins | of ultimate segments obscure. |
of ultimate segments usually evident. |
Sporangia | sessile or subsessile, containing 64 spores, intermixed with abundant farina-producing glands. |
short-stalked, containing 64 spores, not intermixed with farina-producing glands. |
2n | = 58. |
= 58. |
Pellaea bridgesii |
Pellaea cordifolia |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and cliffs, on granitic substrates | Rocky slopes and ledges, usually on volcanic substrates |
Elevation | 1200–3600 m (3900–11800 ft) | 1000–2500 m (3300–8200 ft) |
Distribution |
CA; ID; NV; OR
|
TX; Mexico |
Discussion | The morphology of Pellaea bridgesii is so distinctive that its sectional (and even generic) placement in Pellaea has long been a source of contention. W. H. Wagner Jr. et al. (1983) documented the existence of sterile diploid hybrids (called P. × glaciogena) between P. bridgesii and P. mucronata (see reticulogram), suggesting that P. bridgesii is most closely related to members of sect. Pellaea. In addition to the more obvious characters mentioned above, P. bridgesii is distinguished from other North American species (except P. ternifolia) by its anastomosing veins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The diploid Pellaea cordifolia has often been treated as a variety of the Central American and South American apogamous triploid, P. sagittata. The two taxa are distinguished by a number of qualitative morphologic features (A. R. Smith 1980), and it seems unlikely that they represent cytotypes of a single species. A. F. Tryon (1957) suggested that P. sagittata may have originated through hybridization between P. ovata and P. cordifolia (as P. sagittata var. cordata). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea |
Sibling taxa | ||
Synonyms | Adiantum cordifolium, P. cardiomorpha, P. sagittata var. cordata | |
Name authority | Hooker: Sp. Fil. 2: 238, plate 142b. (1858) | (Sessé & Mociño) A. R. Smith: Amer. Fern J. 70: 26. (1980) |
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