Pellaea breweri |
Pellaea gastonyi |
|
---|---|---|
Brewer's cliff-brake, Brewer's cliffbrake fern |
Gastony's cliff-brake, Gastony's cliffbrake fern |
|
Stems | compact, ascending, stout, 5–10 mm diam.; scales uniformly reddish brown, linear-subulate, 0.1–0.3 mm wide, thin, margins sinuous, nearly entire. |
compact, ascending, stout, 5–10 mm diam.; scales uniformly reddish brown, linear-subulate, 0.1–0.3 mm wide, thin, margins entire to denticulate. |
Leaves | monomorphic, clustered on stem, 2.5–20 cm; croziers sparsely villous. |
somewhat dimorphic, sterile leaves shorter than fertile leaves, clustered on stem, 8–25 cm; croziers villous. |
Petiole | brown, lustrous, rounded adaxially, with prominent articulation lines near base. |
reddish purple to dark brown, lustrous, rounded adaxially, without prominent articulation lines. |
Blade | linear-oblong, pinnate-pinnatifid proximally, 1–4 cm wide; rachis brown proximally, green distally, straight, rounded adaxially, glabrous to sparsely villous. |
elongate-deltate to lanceolate, 2-pinnate proximally, 3–6 cm wide; rachis purple or brown throughout, straight, rounded adaxially, sparsely villous with long, divergent hairs. |
Ultimate segments | lanceolate-deltate, 5–25 mm, herbaceous, glabrous; margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, erose-denticulate; apex obtuse or rounded. |
oblong-lanceolate, 7–30 mm, leathery, sparsely villous abaxially near midrib; margins usually recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, crenulate; apex obtuse to slightly mucronate. |
Pinnae | ascending or perpendicular to rachis, decurrent on rachis, deeply 2-lobed (mitten-shaped) near base of leaf; costae absent. |
ascending or perpendicular to rachis, not decurrent on rachis or obscurely so, usually with 3–7 ultimate segments; costae straight, 2–30 mm, usually shorter than ultimate segments. |
Veins | of ultimate segments evident. |
of ultimate segments obscure. |
Sporangia | sessile or subsessile, containing 64 spores, not intermixed with farina-producing glands. |
long-stalked, containing 32 spores, not intermixed with farina-producing glands. |
2n | = 58. |
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Pellaea breweri |
Pellaea gastonyi |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Cliffs and rocky slopes, on a variety of substrates including granite and limestone | Calcareous cliffs and ledges, usually on limestone |
Elevation | 1600–3800 m (5200–12500 ft) | 100–1500 m (300–4900 ft) |
Distribution |
CA; CO; ID; MT; NV; OR; UT; WA; WY
|
MO; SD; WY; AB; BC; SK |
Discussion | Pellaea breweri is distinguished from other North American taxa (except for some populations of P. glabella) by the presence of prominent articulation lines near the base of the petiole. The leaves are easily detached, and many herbarium specimens consist of separate leaves and stems, the latter covered with petiole bases of approximately equal length. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pellaea gastonyi is an apogamous tetraploid that has originated through repeated hybridization between P. atropurpurea and P. glabella. Isozyme studies (G. J. Gastony 1988) indicate that P. glabella subsp. missouriensis was the diploid parent of plants found in Missouri, whereas diploid P. glabella subsp. occidentalis was involved in the origin of P. gastonyi populations occurring in western North America. Pellaea gastonyi is most often confused with P. atropurpurea, from which it differs in having sparsely villous rachises, smaller ultimate segments, and spores averaging more than 62 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Pellaea | Pteridaceae > Pellaea |
Sibling taxa | ||
Name authority | D. C. Eaton: Proc. Amer. Acad. Arts 6: 555. (1865) | Windham: Contr. Univ. Michigan Herb. 19: 36. (1993) |
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