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Sierra cliff-brake, Sierran cliffbrake

ovate-leaf cliff-brake

Stems

compact, ascending, stout, 5–10 mm diam.;

scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers dark brown to black, thick, margins brown, thin, dentate.

creeping, horizontal, slender, 2–5 mm diam.;

scales mostly bicolored, lanceolate, largest scales 0.3–0.8 mm wide, centers black, thick, margins brown, thin, erose-dentate.

Leaves

monomorphic, clustered on stem, 8–40 cm;

croziers sparsely villous.

monomorphic, clustered or scattered along stem, 15–100 cm;

croziers pubescent, bearing a few scales.

Petiole

dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines.

straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.

Blade

linear-oblong, 2-pinnate proximally, 1–4 cm wide;

rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous.

elongate-deltate, usually 3-pinnate proximally, 5–25 cm wide;

rachis tan throughout, strongly flexuous, rounded or flattened adaxially, usually glabrous.

Ultimate segments

linear, 5–20 mm, leathery, glabrous;

margins on fertile segments strongly revolute, covering more than 1/2 abaxial surface, borders greenish, crenate;

apex mucronate.

lanceolate-deltate, 5–20 mm, leathery, glabrous or sparsely pubescent;

margins recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, entire;

apex obtuse to truncate.

Pinnae

strongly ascending, not decurrent on rachis, usually with 5–11 ultimate segments;

costae straight, 5–20 mm, usually shorter than ultimate segments.

retrorse, projecting downward toward base of leaf, not decurrent on rachis, with 5–40 ultimate segments;

costae strongly flexuous, 25–120 mm, longer than ultimate segments.

Veins

of ultimate segments obscure.

of ultimate segments obscure.

Sporangia

short-stalked, containing 64 spores, intermixed with abundant farina-producing glands.

short-stalked, containing 64 spores, not intermixed with farina-producing glands.

2n

= 58.

Pellaea brachyptera

Pellaea ovata

Phenology Sporulating summer–fall. Sporulating summer–fall.
Habitat Cliffs and rocky slopes, usually on igneous substrates, occasionally on serpentine Rocky slopes and ledges, leaves often supported by associated vegetation, on a variety of substrates including granite and limestone
Elevation 900–2700 m (3000–8900 ft) 300–1700 m (1000–5600 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico; Central America; South America; West Indies in Hispaniola
[WildflowerSearch map]
[BONAP county map]
Discussion

The distinctive Pellaea brachyptera reportedly hybridizes with P. mucronata (A. F. Tryon 1957; D. B. Lellinger 1985); the hybrids are morphologically intermediate plants with malformed spores.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Pellaea ovata in the flora are composed of sexual diploids; an apogamous triploid cytotype predominates south of the United States. I have not seen herbarium specimens to substantiate reports of P. ovata from New Mexico (D. B. Lellinger 1985).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Pellaea Pteridaceae > Pellaea
Sibling taxa
P. andromedifolia, P. atropurpurea, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. intermedia, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
P. andromedifolia, P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. intermedia, P. lyngholmii, P. mucronata, P. ternifolia, P. truncata, P. wrightiana
Synonyms Platyloma brachyptera
Name authority (T. Moore) Baker: in Hooker & Baker, Syn. Fil. ed. 2 477. (1874) (Desvaux) Weatherby: Contr. Gray Herb. 114: 34. 1936 Pteris ovata Desvaux, Mém. Soc. Lin. (1827)
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