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coffee cliffbrake, coffee fern

Stems

creeping, horizontal, slender, 2–4 mm diam.;

scales mostly bicolored, narrowly lanceolate, largest scales 0.3–0.8 mm wide, centers black, thick, margins brown, thin, irregularly dentate.

Leaves

monomorphic, scattered along stem, 10–60 cm;

croziers not conspicuously pubescent, densely scaly.

Petiole

straw-colored, tan, or gray, not lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.

Blade

elongate-deltate, usually 3-pinnate proximally, 3–20 cm wide;

rachis tan throughout, straight to slightly flexuous, rounded or flattened adaxially, glabrous or pubescent.

Ultimate segments

elliptic to ovate, 3–15 mm, somewhat herbaceous, glabrous to sparsely pubescent abaxially;

margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, entire;

apex retuse to rounded.

Pinnae

ascending or perpendicular to rachis, not decurrent on rachis, with 8–50 ultimate segments;

costae usually straight, 15–140 mm, longer than ultimate segments.

Veins

of ultimate segments evident.

Sporangia

short-stalked, containing 64 or 32 spores, not intermixed with farina-producing glands.

2n

= 58;

n = 2n = 87, 116, apogamous.

Pellaea andromedifolia

Phenology Sporulating late spring–summer.
Habitat Rocky slopes and ledges, usually on igneous substrates
Elevation 0–1500 m (0–4900 ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Pellaea andromedifolia comprises three cytotypes: a sexually reproducing diploid, an apogamous triploid, and an apogamous tetraploid. Isozyme studies by G. J. Gastony and L. D. Gottlieb (1985) suggested that the apogamous triploid is an autopolyploid derived from sexual diploid populations. The apogamous tetraploid apparently resulted from hybridization between diploid and triploid individuals. These cytotypes have not been formally recognized as subspecies because their ranges seem to overlap extensively and because the ploidy level of the type collection of P. andromedifolia is not known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2.
Parent taxa Pteridaceae > Pellaea
Sibling taxa
P. atropurpurea, P. brachyptera, P. breweri, P. bridgesii, P. cordifolia, P. gastonyi, P. glabella, P. intermedia, P. lyngholmii, P. mucronata, P. ovata, P. ternifolia, P. truncata, P. wrightiana
Synonyms Pteris andromedifolia, P. andromedifolia var. pubescens
Name authority (Kaulfuss) Fée
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