Pediomelum pentaphyllum |
Pediomelum epipsilum |
|
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Chihuahua scurfpea, small Indian breadroot |
Kane breadroot |
|
Habit | Herbs subacaulescent, to 30 cm, mostly glandular (with obvious blond to dark brown glands) and appressed-spreading pubescent throughout. | Herbs clump-forming, acaulescent or subcaulescent, 3–16(–22) cm, mostly glandular (with obvious blond to dark brown glands) and pubescent throughout. |
Stems | mostly very short-erect, sparsely branched, with leaves clustered at base, rarely with decumbent lateral stems 0–15 cm, with leaves and inflorescences in terminal clusters; pseudoscapes 0–4 cm; cataphylls 7–15 mm (when present), apically emarginate. |
short-erect, unbranched or branched, with 2–5 internodes, sometimes with decumbent lateral stems to 22 cm, strigose to ascending-hairy, leaves appearing clustered basally or on tips of some lateral stems, or dispersed along more elongated stems; pseudoscapes to 4.5 cm; cataphylls to 15 mm. |
Leaves | palmately or pseudopalmately 5(or 6)-foliolate, sometimes with 2 leaflets distal to others; stipules persistent, lanceolate to linear, 9–15 × 5 mm, strigose-tomentose; petiole jointed basally, 60–150 mm, strigose-tomentose; petiolules 1.5–2 mm; leaflet blades lanceolate, rhombic, or slightly oblanceolate, 2–5.5 × (1.5–)2–3.2 cm, base cuneate, margins undulate, apex acute, mucronate, surfaces glandular and pubescent. |
palmately 5(–8)-foliolate; stipules persistent, lanceolate to slightly elliptic, 5–11 × 2.5–4.5 mm, pubescent; petiole jointed basally, 50–80(–100) mm; petiolules 0.5–2 mm; leaflet blades obovate to oblanceolate or ± rhombic, 1.5–2.5(–4) × 1.2–2.5 cm, base cuneate or attenuate, apex broadly acute to rounded, surfaces bicolor, abaxially glandular, cinereous, adaxially bright green, glabrous or sparsely strigose only along base of veins. |
Inflorescences | disjointing in age at peduncle base, globose to long-ovoid; rachis 2–4.5 cm, nodes 6–9, 3 flowers per node; bracts persistent to tardily deciduous, lanceolate to elliptic, often ensiform, 5–7 × 2–3 mm, membranous in age, strigose. |
disjointing in age at peduncle base, subglobose to elongate; rachis 1.4–4 cm, elongating in fruit, nodes 4–9, (1–)3(or 4) flowers per node, internodes relatively short or to 10 mm; bracts persistent to tardily deciduous, ovate to broadly lanceolate, 13–18(–22) × 6–10 mm, apex caudate, pubescent. |
Peduncles | (3–)4–9.5 cm, equal to or shorter than subtending petiole, appressed-pubescent. |
1.4–5 cm, shorter than subtending petiole, pubescent with erect-ascending hairs. |
Pedicels | 2–3 mm. |
2–4 mm. |
Flowers | 14–18 mm; calyx gibbous-campanulate in fruit, 14–16 mm abaxially, 11–12 mm adaxially, glandular, strigose; tube 4–5 mm; abaxial lobe broadly oblanceolate, 10–12 × 3.5–4 mm, adaxial lobes linear-lanceolate, 7–8 × 1–1.5 mm; corolla white to purple, banner white to lavender, paler than other petals, lanceolate to elliptic, 15–17 × 6–7 mm with claw 4–5 mm, wings 15–16 × 3–3.5 mm with claw 6 mm, keel 9–12 × 3 mm with claw 6–7 mm; filaments 8.5–10.5 mm; anthers round-elliptic, 0.3 mm; ovary pubescent apically, style pubescent basally. |
14–19 mm; calyx gibbous-campanulate in fruit, (10–)11–16(–18) mm abaxially, (9–)10–15(–16) mm adaxially, tube glandular, pubescent; tube 5–6(–8) mm; lobes linear-lanceolate to elliptic, abaxial 6.5–10 × 2–3 mm, adaxial 6–9 × 1–1.5 mm, glandular or eglandular; corolla purple, banner sometimes paler, oblanceolate, 14–19 × 5–7 mm with claw 6–9 mm, wings 9–18 × 2–3 mm with claw 6–9 mm, keel (10–)12–16 × 2–3 mm with claw 7–9 mm; filaments 11–16 mm; anthers elliptic, 0.3 mm; ovary pubescent or only apically, style pubescent basally. |
Legumes | ellipsoid-oblanceoloid, 7–8 × 4–5 mm, eglandular, pubescent, beak broad, flat, 10–15 mm, barely exserted beyond calyx. |
ovoid, 6–9 × 4–4.5 mm, eglandular, pubescent, beak 5–8 mm, ± equal to calyx. |
Seed | gray-green to brown, oblong-reniform, 5–6 × 3–4 mm, rugose. |
brown, reniform, 4–5 × 3 mm, shiny. |
Pediomelum pentaphyllum |
Pediomelum epipsilum |
|
Phenology | Flowering spring–early summer. | Flowering late spring–summer. |
Habitat | Desert grasslands. | Rocky to clay soils, pine or juniper woodlands, desert shrub communities. |
Elevation | 1500–2000 m. (4900–6600 ft.) | 1600–1700 m. (5200–5600 ft.) |
Distribution |
AZ; NM; TX; Mexico (Chihuahua) |
AZ; UT |
Discussion | A history of the confusing nomenclature of Pediomelum pentaphyllum was given by P. Tonne (2000). The name was misapplied by P. A. Rydberg (1919–1920); the plant that Rydberg described as P. pentaphyllum was actually P. palmeri; the plant denoted here as P. pentaphyllum was called P. trinervatum Rydberg. J. W. Grimes (1990), before completion of his monograph, erroneously annotated several P. pentaphyllum specimens as P. trinervatum. Pediomelum pentaphyllum is rare, with only a few confirmed populations in Arizona (Cochise and Graham counties) and New Mexico (Hidalgo County); it was also collected in Texas (Presidio County), and Chihuahua, Mexico (P. Tonne 2000), but may be extirpated in those states. Pediomelum pentaphyllum is said to have been used by Native Americans and indigenous Mexicans as a fever reducer, the use of which may have contributed historically to its spread through cultivation. It is uncertain as to why P. pentaphyllum has declined, but causes may include over-collecting, grazing, and herbicide application (WildEarth Guardians, https://pdf.wildearthguardians.org/site/DocServer/petition_scurfpea.pdf?docID=624&AddInterest=1103). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pediomelum epipsilum is known from Coconino and Mohave counties in Arizona and Kane County in Utah. It has been variously treated at specific and varietal rank, but bract and leaflet morphology, as well as phylogenetic data (A. N. Egan and K. A. Crandall 2008, 2008b), support recognition of the taxon at the rank of species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Psoralea pentaphylla | Psoralea epipsila, P. megalanthum var. epipsilum |
Name authority | (Linnaeus) Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 23. (1919) | (Barneby) S. L. Welsh: Great Basin Naturalist 46: 257. (1986) |
Web links |