Pediomelum pariense |
Pediomelum subacaule |
|
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Paria Plateau Indian breadroot, Paria River Indian breadroot |
Nashville breadroot, white-rim scurfpea |
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Habit | Herbs acaulescent or subacaulescent, 2–9 cm, mostly glandular and strigose throughout. | Herbs acaulescent or subacaulescent, to 20 cm, eglandular except leaves or rarely stipules and bracts, pubescent. |
Stems | short-erect, rarely branched (when branched then branches subtended by cataphylls), leaves clustered; pseudoscapes to 2.5 cm; cataphylls 6–11 mm, glabrous. |
absent, leaves clustered; pseudoscapes erect, 3–10 cm, usually 1, sometimes to 6, rarely branched proximally; cataphylls 10–20 mm (when present). |
Leaves | palmately 3(–5)-foliolate; stipules persistent, obovate to oval, 4–10 × 2.5–6 mm, glabrous or sparsely pubescent; petiole not enlarged but jointed basally, slightly canaliculate, 13–70 mm; petiolules 0.5–1 mm; leaflet blades obovate, 0.9–2.5 × 0.7–2.2 cm, base cuneate or attenuate, apex rounded or emarginate, surfaces abaxially gray-green and evenly strigose, adaxially yellow-green and strongly white-strigose along veins. |
palmately 5–7-foliolate; stipules often fragmented on plant, partly connate, ovate, 11–24 × 11–16 mm, scarious, usually eglandular, sparsely pubescent; petiole jointed basally, 40–150(–190) mm, shorter than peduncle, hirsute-villous; petiolules 1–2.5 mm; leaflet blades oblanceolate to elliptic, 1.5–4.5(–6) × 0.5–1.3(–1.8) cm, base cuneate, apex acuminate to obtuse, surfaces glandular, abaxially pubescent, adaxially sparsely so, usually along margins. |
Inflorescences | disjointing in age at peduncle base, globose; rachis 0.5–8.5 cm, elongating through fruiting, nodes (1 or)2–6, 3 flowers per node, internodes 1–3 mm; bracts tardily deciduous or persistent, oblanceolate to elliptic, 4–8 × 3–4 mm, apex abruptly acuminate, short-pubescent. |
disjointing in age at peduncle base, long-ovoid; rachis 2.5–9.5 cm, elongating through fruiting, nodes 4–11(–15), 3 flowers per node, internodes 1–5 mm, elongating to 25 mm in fruit; bracts persistent, orbiculate to lanceolate, 7–14 × 2–7 mm, margins pubescent, apex sometimes caudate or emarginate, becoming papery and veined in age. |
Peduncles | 2–4.5 cm, shorter than subtending petiole, pubescent. |
4–14 cm (8–19 cm in fruit), longer than subtending petiole, setose. |
Pedicels | 2–5 mm. |
1–2 mm. |
Flowers | 8–12.5 mm; calyx broadly gibbous-campanulate in fruit, 10–11.5 mm abaxially, 8–8.5 mm adaxially, eglandular or sparsely glandular, pubescent; tube 3.3–4.5 mm; lobes lanceolate or oblanceolate, abaxial 4–6.5 × 2–3.5 mm, adaxial 3–5 × 1–1.5 mm; corolla ochroleucous, wings and keel suffused purple or purple, banner broadly orbiculate to obovate, (8.5–)10–14 × 5.5–7.5 mm with claw 2.5–3 mm, wings 9–11 × 2.5–3 mm with claw 3.5–4 mm, keel 8–8.5 × 2.5–3 mm with claw 3–4 mm; filaments 7–8 mm; anthers broadly elliptic, 0.4 mm; ovary pubescent apically, style pubescent basally. |
10–15(–22) mm; calyx weakly gibbous-campanulate in fruit, 6–10 mm abaxially, 5–8 mm adaxially, glabrate and eglandular on gibbous portion, sparsely setose and glandular distally; tube stramineous, 4.5–6 mm; lobes triangular, abaxial 2–4 × 2 mm, adaxial 1–2 × 1 mm; corolla dark blue to purple, banner oblanceolate to oval, 13–23 × 4–5 mm with claw 5–6 mm, wings 12–18 × 2–3 mm with claw 5–7 mm, keel 10–15 × 2–3 mm with claw 2–3 mm; filaments 10–12 mm; anthers obovate, 0.3 mm; ovary pubescent apically, style pubescent basally. |
Legumes | ellipsoid, 6–7 × 4–4.5 mm, eglandular, strigose, beak 3–4 mm, not exserted beyond calyx. |
globose to ovoid, 6–7 × 4–5 mm, eglandular, sparsely pubescent apically, beak triangular, 4–6 mm, exserted beyond calyx. |
Seed | gray-green to brown, obovoid, 4.5–5 × 2.5–3 mm, shiny. |
gray-brown to red-brown, obovoid-reniform, 5–6 × 3–4 mm. |
2n | = 22. |
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Pediomelum pariense |
Pediomelum subacaule |
|
Phenology | Flowering spring–summer. | Flowering spring. |
Habitat | Barren outcrops in open pine-juniper woodlands. | Calcareous soils, cedar glades. |
Elevation | 1700–2500 m. (5600–8200 ft.) | 200–1500 m. (700–4900 ft.) |
Distribution |
UT |
AL; GA; TN
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Discussion | Pediomelum pariense is known from Garfield, Kane, and Washington counties. It much prefers open, barren rocky soils, particularly limestone and is considered rare, with fewer than 20 populations known. A few known populations inhabiting the high-elevation escarpment ridges of Bryce Canyon National Park are assured protection, but the bulk of populations exist on the lower steps of the Grand Staircase-Escalante area, areas that may become targets for mining and oil developments. In 2005 and 2006, considerable effort was spent to seek out new populations but without success. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pediomelum subacaule is known from Colbert and Franklin counties in Alabama, Catoosa County in Georgia, and Davidson, Maury, Rutherford, and Wilson counties in Tennessee. Pediomelum subacaule is quite distinct and disjunct from its fellow members of subg. Disarticulatum, this evidenced by its large, partly connate stipules and short, broad calyx lobes, and by being the only species in the subgenus east of the Mississippi River. Pediomelum subacaule is restricted to limestone soil in open cedar glades. Although P. subacaule is well established in the protected cedar glades of Tennessee, its historical range is shrinking due to habitat loss. Several historical sites that were, or still are, active limestone quarries have no remaining evidence of P. subacaule, particularly in Alabama and Georgia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Psoralea pariensis | Psoralea subacaulis, Lotodes subacaulis |
Name authority | (S. L. Welsh & N. D. Atwood) J. W. Grimes: Brittonia 38: 185. (1986) | (Torrey & A. Gray) Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 20. (1919) |
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