Pediocactus nigrispinus |
Pediocactus knowltonii |
|
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black-spine snowball cactus, Columbia Plateau cactus, dark-spine ball cactus, snowball cactus |
Knowlton's miniature cactus, Knowlton's minute cactus |
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Habit | Plants typically branched. | Plants branched or unbranched. |
Stems | depressed-ovoid to elongate-ovoid, 5–30 × 5–15 cm; areoles oval, villous. |
globular to short cylindric, 0.7–5.5 × 1–3 cm; areoles circular, villous. |
Spines | smooth, hard and rigid, distinguishable as radial and central; radial spines 10–30 per areole, spreading at right angles to tubercles, nearly straight, white to dull reddish brown, 8–20 mm; central spines 6–12 per areole, widely spreading or nearly erect, reddish brown to nearly black, rigid, straight or slightly curved, base yellow or cream, 15–35 mm, less than 1 mm diam. at base. |
smooth, relatively hard, all radial, mostly 18–26 per areole, spreading, recurved, or somewhat pectinate, reddish tan, pink, or white, 1–1.5 mm, canescent. |
Flowers | 1–3.5 × 2.5–5 cm; scales and outer tepals of flower tube minutely toothed, laciniate, or entire and undulate; outer tepals with greenish brown midstripes, oblong-cuneate, 12–25 × 4.5–9 mm; inner tepals white, pink, magenta, yellow, or yellow-green, 19–27 × 5–10 mm. |
1–3.5 × 1–2.5 cm; scales and outer tepals essentially entire, often undulate; outer tepals with brownish midstripes to 1.5 mm wide, 4–17 × 4–6 mm; inner tepals pink, oblanceolate, 8–25 × 3–8 mm. |
Fruits | green tinged with red, drying reddish brown, short cylindric, 6–11 × 5–10 mm. |
green, drying reddish tan, turbinate,4 × 3 mm. |
Seeds | gray to black, 2–3 × 1.5–2 mm, papillate but not rugose. |
black, 1.5 × 1–1.2 mm, papillate but not rugose. |
Pediocactus nigrispinus |
Pediocactus knowltonii |
|
Phenology | Flowering spring. | Flowering spring. |
Habitat | Great Basin desert scrub, sagebrush, grasslands, coniferous forests | Gravel pavements in pinyon-juniper woodlands with mixed sagebrush |
Elevation | 400-2000 m (1300-6600 ft) | 2000 m (6600 ft) |
Distribution |
ID; OR; WA
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NM |
Discussion | No known morphologic character supports the taxonomic recognition of infraspecific taxa within Pediocactus nigrispinus. Characteristics used to distinguish the three described subspecies almost completely overlap. Pediocactus nigrispinus has been referred to P. simpsonii var. robustior (J. M. Coulter) L. D. Benson, which remains well within the range of variation for P. simpsonii. An unpublished study by J. M. Porter et al. of noncoding chloroplast DNA sequences shows P. simpsonii is less closely related to P. nigripsinus than to P. knowltonii, P. winkleri, and P. despainii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Pediocactus knowltonii is the smallest and rarest member of the genus. The species is very closely related to P. simpsonii; chloroplast DNA sequence analysis provides support that it represents a recent developmental mutation within P. simpsonii (J. M. Porter et al. unpubl.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4. | FNA vol. 4. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | P. simpsonii var. nigrispinus, P. nigrispinus var. beastonii, P. nigrispinus subsp. beastonii, P. nigrispinus subsp. puebloensis | P. bradyi var. knowltonii, P. simpsonii var. knowltonii |
Name authority | (Hochstätter) Hochstätter: Succulenta (Netherlands) 71: 99. (1992) | L. D. Benson: Cact. Succ. J. (Los Angeles) 32: 193. (1960) |
Web links |