Pedicularis sudetica
Pedicularis capitata
fernweed, Oeder's lousewort, sudetan lousewort, sudeten lousewort, sudetic lousewort
capitate lousewort, pédiculaire capitée
basal 1–20, blade elliptic to lanceolate, 10–110 x 3–26 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous;
cauline 0–5, blade lanceolate to elliptic, 20–90 x 2–20 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, 1- or 2-serrate, surfaces glabrous, some hairs along veins on abaxial surface.
basal 2 or 3, blade ovate or elliptic, 10–40 x 5–20 mm, 2(or 3)-pinnatifid, margins of adjacent lobes nonoverlapping or extensively overlapping distally, serrate, surfaces glabrous or hirsute;
cauline 0–2, blade elliptic, 5–25 x 5–10 mm, 2-pinnatifid, margins of adjacent lobes nonoverlapping or extensively overlapping distally, serrate, surfaces scattered hispid.
simple, 1–4, exceeding basal leaves, each 10–50-flowered;
bracts linear to subulate or trullate, 2–15 x 1–4 mm, undivided with or without long auricles, or 1-pinnatifid, margins entire, serrate, or serrulate, surfaces glabrous, white- or yellowish white-lanate, or sparsely pilose.
simple, capitate, 1 or 2, exceeding basal leaves, each 2–8-flowered;
bracts subulate to narrowly lanceolate, 10–20(–50) x 6–8 mm, undivided proximally, undivided or 1- or 2-pinnatifid distally, proximal margins entire, distal serrate, surfaces sparsely tomentose.
1–2.5 mm.
2–3 mm.
calyx 7–13 mm, glabrous, white-lanate, yellowish white-lanate, or sparsely pilose, lobes 5, subulate or triangular, 1.5–5 mm, apex entire, crenulate, or serrulate, glabrous, sometimes ciliate;
corolla 16–21 mm, tube pink, purple, or magenta, 9–11 mm;
galea purple, magenta, or bicolored, 7–12 mm, beakless, margins entire medially, 1-toothed or entire distally, apex arching over abaxial lip;
abaxial lip white or pink with purple spots, purple, or magenta, 4–8 mm.
calyx 10–15 mm, hirsute, lobes 5, triangular to spatulate, 4–6 mm, apex serrate, glabrous;
corolla 19–40 mm, tube light yellow, sometimes cream to pink, 5–20 mm;
galea light yellow, sometimes cream to pink, apically sometimes diffuse purple, 12–20 mm, beakless, margins entire medially and distally, apex arching beyond abaxial lip;
abaxial lip light yellow, sometimes cream to pink, 10–15 mm.
= 16 (Asia).
Pedicularis sudetica
Pedicularis capitata
Subspecies 6 (5 in the flora).
Pedicularis sudetica is a difficult complex. Hultén employed the degree of lobing on subtending floral bracts, length of corolla tubes, and inflorescence vestiture in recognizing eight infraspecific taxa. U. Molau and D. F. Murray (1996) emphasized presence or absence of spots on the abaxial lip, inflorescence vestiture, length of petioles and calyx, and ecological features to define four species, subsuming several of Hultén’s other subspecific taxa into these species. According to Molau and Murray, P. sudetica in the narrow sense is a morphologically distinct, disjunct taxon endemic to the Sudeten Mountains of central Europe, but part of the broader circumscription by Hultén (1961, 1964), which is treated here as the sixth subspecies.
U. Molau and D. F. Murray (1996) did not include subsp. scopulorum in their analysis, noting only that teeth are virtually absent on the galea of Pedicularis scopulorum. Presence or absence of apical teeth on the margins of the galea often distinguishes species in other Pedicularis taxa. Because teeth are sometimes present in this taxon, it is treated here as one of the five North American subspecies in the broad sense of P. sudetica. A recent molecular study confirms its close relationship to other members of the complex (B. W. Robart et al. 2015).
When comparing the two alternative taxonomies, the treatment by U. Molau and D. F. Murray (1996) is easier to apply. Close inspection of specimens identified as these species often reveals combinations of traits attributable to more than one taxon; Molau and Murray reported finding hybridization common between Pedicularis albolabiata and P. arctoeuropaea, and P. albolabiata and P. pacifica where habitats overlap, indicating that reproductive isolation is not complete. Recognition of these taxa as varieties may be more appropriate considering the broad geographic overlap where the few distinguishing features tend to intergrade. They are treated here, however, as subspecies. The following key is modified from Molau and Murray.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Inflorescences of Pedicularis capitata display no more than five clustered flowers; the long, vertical corolla tube and curved galea that arches over the appressed, broad lobes of the abaxial lip are distinctive. The tip of the galea can be the same color as the corolla tube and abaxial lip but sometimes is a diffuse pink to purple and not distinctly bicolored as in the galeas of P. flammea or P. oederi, with which it is often confused.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Calyx lobe margins entire distally. | → 2 |
2. Calyx lobes subulate; Alberta, British Columbia, Manitoba, Northwest Territories, Nunavut, Ontario, Yukon, Alaska. | subsp. interior |
2. Calyx lobes triangular; Colorado, New Mexico, Wyoming. | subsp. scopulorum |
1. Calyx lobe margins crenulate or serrulate distally. | → 3 |
3. Corollas: abaxial lips purple-spotted. | → 4 |
4. Corollas: galeas bicolored; abaxial lips white, rarely pink; calyces and bracts glabrous or white-lanate. | subsp. albolabiata |
4. Corollas: galeas not bicolored; abaxial lips magenta; calyces and bracts yellowish white-lanate, rarely glabrous. | subsp. arctoeuropaea |
3. Corollas: abaxial lips not purple-spotted. | → 5 |
5. Calyces and bracts white-lanate, calyx lobe margins entire or serrulate distally. | subsp. interior |
5. Calyces and bracts glabrous or sparsely pilose, calyx lobe margins serrulate distally. | subsp. pacifica |
BC
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
