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swamp lousewort

Howell lousewort, Howell's lousewort, Howell's pedicularis

Habit Plants 20–100 cm. Plants 15–40 cm.
Leaves

basal 0;

cauline 10–30, blade lanceolate, 20–100 x 10–30 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, 2-serrate, surfaces hispid.

basal 1–4, blade elliptic to lanceolate, 20–60 x 15–40 mm, undivided or 1-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous;

cauline 10–12, blade ovate to lanceolate, 25–60 x 15–40 mm, undivided or 1-pinnatifid, sometimes auricled, margins of adjacent lobes nonoverlapping or slightly overlapping distally, entire or serrate to crenate, surfaces glabrous or scattered woolly along main vein.

Racemes

paniculate, 1, each 10–20-flowered;

bracts lanceolate to trullate, 5–10 x 3–5 mm, undivided or 1-pinnatifid and 1- or 2-auricled, proximal margins entire, distal 1- or 2-serrate, surfaces glabrous or hispid.

simple, 1–8, exceeding basal leaves, each 15–40-flowered;

bracts trullate to cordate, 6–8 x 4–8 mm, undivided, proximal margins entire, distal entire, surfaces glabrous or tomentose.

Pedicels

1–1.5 mm.

2.5–3 mm.

Flowers

calyx 7–12 mm, glabrous or hispid, lobes 2, trullate, ovate, elliptic, or triangular, 2.5–3.5 mm, apex serrate, glabrous, sometimes ciliate;

corolla 16–22 mm, tube white, cream, or light yellow, 8–12 mm;

galea white, cream, or light yellow, 8–12 mm, beaked, beak straight, 0.5–2.5 mm, margins entire medially and distally, apex extending over abaxial lip;

abaxial lip white, cream, or light yellow, 7–10 mm.

calyx 6–6.5 mm, tomentose, lobes 5, triangular, 1.5–2 mm, apex entire, ciliate;

corolla 10–13 mm, tube white, 6–8 mm;

galea white, apically sometimes tinged with red to violet, 3.5–5 mm, beaked, beak straight, 1–2 mm, margins entire medially and distally, apex extending beyond abaxial lip;

abaxial lip white, 1.5–2 mm.

2n

= 16.

Pedicularis lanceolata

Pedicularis howellii

Phenology Flowering Aug–Oct. Flowering Jun–Aug.
Habitat Wet meadows, fens, springs, moist prairies, swamps. Alpine forest clearings and edges.
Elevation 10–1100 m. (0–3600 ft.) 1100–2000.
Distribution
from FNA
AR; CT; DE; GA; IA; IL; IN; KY; MA; MD; MI; MN; MO; NC; ND; NE; NJ; NY; OH; PA; SD; TN; VA; WI; WV; MB; ON
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[BONAP county map]
from FNA
CA; OR
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[BONAP county map]
Discussion

The long corolla tubes of Pedicularis lanceolata are uncharacteristically nectarless, and only late season pollen-foraging worker bumblebees pollinate this species (L. W. Macior 1969). The uniquely hinged abaxial lip covering the opening of the galea is an adaptation to allow only worker bumblebees access to the anthers, as they must learn to push it aside during foraging.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pedicularis howellii is found in the Siskiyou Mountains along the California/Oregon border; it has undivided distal leaves. The division of the proximal leaves into irregular and asymmetric lobes, sometimes appearing auricled, is a unique feature of this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 525. FNA vol. 17, p. 523.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Name authority Michaux: Fl. Bor.-Amer. 2: 18. (1803) A. Gray: Proc. Amer. Acad. Arts 20: 307. (1885)
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