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swamp lousewort

attol lousewort, attoll lousewort, elephant snouts, little elephant head, little elephant's head, woolly Mammoth

Habit Plants 20–100 cm. Plants 15–78 cm.
Leaves

basal 0;

cauline 10–30, blade lanceolate, 20–100 x 10–30 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, 2-serrate, surfaces hispid.

basal 5–25, blade elliptic, 60–150(or 200–250) x 3–23 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous or scattered glands;

cauline 2–20, blade elliptic, 5–50(–100) x 1–5 mm, undivided or 1(or 2)-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous.

Racemes

paniculate, 1, each 10–20-flowered;

bracts lanceolate to trullate, 5–10 x 3–5 mm, undivided or 1-pinnatifid and 1- or 2-auricled, proximal margins entire, distal 1- or 2-serrate, surfaces glabrous or hispid.

simple, 1–3, exceeding basal leaves, each 10–50-flowered;

bracts lanceolate to triangular, 5–10 x 3–10 mm, pinnatifid, margins entire, surfaces glabrous or tomentose.

Pedicels

1–1.5 mm.

1.2–1.6 mm.

Flowers

calyx 7–12 mm, glabrous or hispid, lobes 2, trullate, ovate, elliptic, or triangular, 2.5–3.5 mm, apex serrate, glabrous, sometimes ciliate;

corolla 16–22 mm, tube white, cream, or light yellow, 8–12 mm;

galea white, cream, or light yellow, 8–12 mm, beaked, beak straight, 0.5–2.5 mm, margins entire medially and distally, apex extending over abaxial lip;

abaxial lip white, cream, or light yellow, 7–10 mm.

calyx 4–5 mm, glabrous or tomentose, lobes 5, triangular, 2–2.5 mm, apex entire, glabrous;

corolla 6–8 mm, tube pink, rarely white, 3–6 mm;

galea white or pink with 2 purple spots or stripes, 1–2 mm, beaked, beak coiled, 3–6 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly vertical;

abaxial lip pendulous, white or pink with purple stripe, 4–5.5 mm.

2n

= 16.

= 16.

Pedicularis lanceolata

Pedicularis attollens

Phenology Flowering Aug–Oct.
Habitat Wet meadows, fens, springs, moist prairies, swamps.
Elevation 10–1100 m. (0–3600 ft.)
Distribution
from FNA
AR; CT; DE; GA; IA; IL; IN; KY; MA; MD; MI; MN; MO; NC; ND; NE; NJ; NY; OH; PA; SD; TN; VA; WI; WV; MB; ON
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from FNA
CA; NV; OR
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[BONAP county map]
Discussion

The long corolla tubes of Pedicularis lanceolata are uncharacteristically nectarless, and only late season pollen-foraging worker bumblebees pollinate this species (L. W. Macior 1969). The uniquely hinged abaxial lip covering the opening of the galea is an adaptation to allow only worker bumblebees access to the anthers, as they must learn to push it aside during foraging.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

The flowers of Pedicularis attollens, like those of P. groenlandica, resemble an elephant’s head, and A. Heller placed them both in Elephantella. The short, upturned beak, in contrast to the long, more horizontal downturned beak of P. groenlandica, is a distinguishing feature of P. attollens. Whereas P. groenlandica occurs across much of western and arctic North America, P. attollens is found primarily in the Cascade Range of central and southern Oregon and the Sierra Nevada of California and Nevada. It is also reported from the Klamath Range to the west and the White and Sweetwater mountains and the Warner Range to the east of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 1-pinnatifid.
subsp. attollens
1. Basal leaves 2-pinnatifid.
subsp. protogyna
Source FNA vol. 17, p. 525. FNA vol. 17, p. 514.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. attollens subsp. attollens, P. attollens subsp. protogyna
Synonyms Elephantella attollens
Name authority Michaux: Fl. Bor.-Amer. 2: 18. (1803) A. Gray: Proc. Amer. Acad. Arts 7: 384. (1868)
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