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Pedicularis densiflora

Indian warrior, warrior's plume

Parry's lousewort

coil-beak lousewort, coiled lousewort, coiled pedicularis, curve-beak lousewort, curved-beak lousewort, white coiled-beak lousewort

Habit Plants 10–50 cm. Plants 7–65 cm. Plants 10–50 cm.
Leaves

basal 1–10, blade lanceolate, 30–200 x 20–70 mm, 2(or 3)-pinnatifid, margins of adjacent lobes nonoverlapping or extensively overlapping distally, 1-serrate, surfaces glabrous, hispid, or downy;

cauline 4–20, blade lanceolate, 15–250 x 5–100 mm, 2-pinnatifid, margins of adjacent lobes nonoverlapping or extensively overlapping distally, serrate, surfaces glabrous, hispid, or downy.

basal 4–20, blade elliptic or lanceolate to oblanceolate, 10–70 x 3–15 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, 1- or 2-serrate, surfaces glabrous;

cauline 0–20, blade lanceolate, 10–50 x 2–10 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous or tomentose.

basal 3–10, blade lanceolate, 30–100 x 5–30 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, entire or serrate, surfaces glabrous;

cauline 2–10, blade elliptic or triangular to lanceolate, 5–70 x 10–25 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, entire or serrate, surfaces glabrous.

Racemes

simple, 1–5, exceeding basal leaves, each 10–50-flowered;

bracts lanceolate to trullate, 10–35 x 3–5 mm, undivided or 1-pinnatifid, proximal margins entire, distal 1- or 2-serrate, surfaces glabrous.

simple, 1–10, exceeding basal leaves, each 5–50-flowered;

bracts subulate to trullate, 10–25 x 8–10 mm, undivided or +/- lobed, proximal margins entire, distal serrate or crenate, surfaces glabrous or tomentose.

simple, 1–6, exceeding basal leaves, each 12–40-flowered;

bracts trullate to obtrullate, 5–18 x 2–15 mm, pinnatifid, margins entire, surfaces glabrous.

Pedicels

2–4 mm.

2–3 mm.

1.5–5.5 mm.

Flowers

calyx 9–18 mm, downy to tomentose, lobes 5, triangular, 3–4 mm, apex entire, ciliate;

corolla 23–43 mm, tube dark red, purple, or orange-yellow, rarely white, 8–18 mm;

galea dark red, purple, or orange-yellow, rarely white, 15–25 mm, beakless, margins entire medially and distally, apex straight;

abaxial lip dark red, purple, or orange-yellow, rarely white, 8–15 mm.

calyx 6–10 mm, glabrous or tomentose, lobes 5, triangular to deltate, 1–3 mm, apex entire, glabrous;

corolla 14–22 mm, tube white, yellowish, or light purple to purple, 7–15 mm;

galea white, yellowish, or light purple to purple, 7–10 mm, beaked, beak straight, 5–8 mm, margins entire medially and distally, apex extending beyond abaxial lip;

abaxial lip white, yellowish, or light purple to purple, 4–9 mm.

calyx 5–9 mm, glabrous, lobes 5, triangular, 1–3 mm, apex entire, glabrous;

corolla 7–13 mm, tube white or cream to yellowish or pink to pinkish purple, 4–7 mm;

galea white or cream to yellowish with purple spots at base, or pink to pinkish purple, 2–5.5 mm, beaked, beak coiled, 5–9 mm, base curving, margins entire medially and distally, apex surrounded by abaxial lip, axis of coil nearly vertical;

abaxial lip surrounding beak, white or cream to yellowish, or pink to pinkish purple, 5–8 mm.

2n

= 16.

= 16.

Pedicularis densiflora

Pedicularis parryi

Pedicularis contorta

Phenology Flowering Feb–May.
Habitat Mixed coniferous forests.
Elevation 30–3100 m. (100–10200 ft.)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MT; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Scarlet corollas with an undomed, toothless galea and two- or three-pinnatifid leaves are diagnostic of Pedicularis densiflora. This species occurs in forested subalpine regions of southern Oregon, western slopes of the Sierra Nevada, and the Coast Ranges of California south to Baja California. Herbarium records indicate northern populations of P. densiflora occur at higher elevations than do more southern populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Pedicularis parryi is characterized by a domed galea with a conical beak and light green calyces and floral bracts with dark purple stripes. The beaked galea extends well beyond the expanded lobes of the abaxial lip.

Pedicularis parryi is an alpine complex. Corolla color, number of cauline leaves, and bract margin variation are important features differentiating infraspecific taxa within P. parryi (G. D. Carr 1971, from which the following key is adapted).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

The coiled beak that extends directly downward from the top of the galea is distinctive in Pedicularis contorta. Seen from the side, the large, upward-expanded, lateral lobes of the abaxial lip surround the beak, concealing it from view. The flowers of P. contorta are very similar to those of P. racemosa; however, the leaves of P. racemosa are undivided, the beak curves down and to one side, and the inflorescence often branches, forming a panicle of short racemes with long internodes between flowers.

Flower and calyx color and width of the subtending floral bracts delimit three varieties in Pedicularis contorta; corolla color is difficult to discern in herbarium specimens. The following key is adapted from R. N. Reese (1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corolla tubes +/- purple; calyces tomentose.
P. parryi subsp. purpurea
1. Corolla tubes white or yellowish; calyces glabrous.
→ 2
2. Cauline leaves (0 or)1–6; bract margins entire or +/- crenate.
P. parryi subsp. parryi
2. Cauline leaves 5–20; bract margins +/- serrate.
P. parryi subsp. mogollonica
1. Corolla tubes, galeas, and abaxial lips white or cream to yellowish, galeas with purple spots at base; calyces pale green.
P. contorta var. contorta
1. Corolla tubes, galeas, and abaxial lips pink to pinkish purple; calyces green or reddish with purple spots.
→ 2
2. Calyces green; bracts 5–8 mm wide, proximal margins ciliate.
P. contorta var. ctenophora
2. Calyces reddish with purple spots; bracts 2–4 mm wide, proximal margins glabrous.
P. contorta var. rubicunda
Source FNA vol. 17, p. 521. FNA vol. 17, p. 528. FNA vol. 17, p. 519.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. parryi subsp. mogollonica, P. parryi subsp. parryi, P. parryi subsp. purpurea
P. contorta var. contorta, P. contorta var. ctenophora, P. contorta var. rubicunda
Name authority Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 110. (1838) A. Gray: Amer. J. Sci. Arts, ser. 2, 34: 250. (1862) Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 108. (1838)
Web links 
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