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fern-leaf lousewort, fernleaf pedicularis

bract lousewort, fernleaf, towering lousewort, wood-betony

Habit Plants 10–50 cm. Plants 20–80 cm.
Leaves

basal 2–10, blade elliptic to lanceolate, 20–90 x 5–15 mm, 2(or 3)-pinnatifid, margins of adjacent lobes nonoverlapping to extensively overlapping distally, serrate, surfaces glabrous;

cauline 2–8, blade triangular to lanceolate, 20–120 x 5–20 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or extensively overlapping distally, serrate, surfaces glabrous or unevenly hispid to tomentose.

basal 0–10, blade lanceolate, 20–120 x 10–60 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, 1- or 2-serrate, surfaces glabrous;

cauline 4–10, blade lanceolate, 10–270 x 8–150 mm, undivided or 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate to 2-serrate, surfaces glabrous or scattered glandular.

Racemes

simple, 1–2, exceeding basal leaves, each 10–40-flowered;

bracts trullate to obtrullate or subulate to trullate, 10–25 x 2–5 mm, undivided or 1- or 2-auricled, sometimes 1-pinnatifid, proximal margins entire, distal entire or serrate, surfaces tomentose.

simple, 1–4, exceeding basal leaves, each 15–75-flowered;

bracts lanceolate or subulate to trullate, 10–20 x 2–10 mm, undivided, proximal margins entire, distal entire or serrate, surfaces glabrous, hispid, or tomentose.

Pedicels

1–3 mm.

0.5–1 mm.

Flowers

calyx 8–12 mm, tomentose, lobes 5, narrowly triangular, 3–4 mm, apex entire, ciliate;

corolla 20–26 mm, tube red or pink, 13–15 mm;

galea red or pink, 7–11 mm, beakless, margins entire medially, 1-toothed distally, apex arching over abaxial lip;

abaxial lip red or pink, 6–7.5 mm.

calyx 7–15 mm, glabrous or tomentose, lobes 5, triangular or filiform, 1–10 mm, apex entire, glabrous or ciliate;

corolla 14–27 mm, tube yellow or dark blood red, 6–12 mm;

galea yellow to yellow tinged with red, purple tinged with yellow, or dark blood red, 6–15 mm, beakless or beaked, beak straight, 0.8–2.5 mm, margins entire medially and distally, apex arching over abaxial lip;

abaxial lip expanded, yellow, yellow tinged with purple, or dark blood red, 4.5–6.5 mm.

2n

= 32.

Pedicularis cystopteridifolia

Pedicularis bracteosa

Phenology Flowering May–Aug.
Habitat Rocky alpine tundras, meadows.
Elevation 2100–3100 m. (6900–10200 ft.)
Distribution
from FNA
MT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Pedicularis cystopteridifolia occurs only in the Rocky Mountains of Montana and Wyoming. As the specific epithet implies, the leaves strongly resemble those of the fern Cystopteris fragilis. Although not sympatric, this species could easily be misidentified as P. sudetica subsp. scopulorum, which has flowers of a similar shape and color, and leaves that are also two-pinnatifid. The secondary leaf lobes of P. cystopteridifolia, however, are much larger, longer, more deeply incised, and more heavily toothed, making them appear more finely dissected than the linear to deltate secondary and smaller toothed lobes of P. sudetica. Many of the adjacent leaf lobes of P. cystopteridifolia also overlap, whereas the lobes of P. sudetica are more widely spaced and therefore not overlapping. The galea of P. cystopteridifolia is also more highly domed and broader, and the leaves are a paler shade of green in contrast to the dark green leaves of P. sudetica subsp. scopulorum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 8 (8 in the flora).

Pedicularis bracteosa is found in subalpine habitats across much of western North America, occurring throughout the Rocky Mountains from central British Columbia and Alberta to northern New Mexico, as well as the Coast Range south to northern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Galeas beaked.
→ 2
2. Calyces tomentose.
var. canbyi
2. Calyces glabrous.
var. siifolia
1. Galeas beakless.
→ 3
3. Calyx lobes filiform.
→ 4
4. Corollas: tubes yellow, galeas yellow to yellow tinged with red, or purple tinged with yellow, abaxial lips yellow to yellow tinged with purple.
var. bracteosa
4. Corollas: tubes, galeas, and abaxial lips dark blood red.
var. atrosanguinea
3. Calyx lobes triangular.
→ 5
5. Galeas 10–15 mm.
var. paysoniana
5. Galeas 7–11 mm.
→ 6
6. Galea apices acute; British Colombia, Idaho, Washington.
var. latifolia
6. Galea apices obtuse; Blue and Wallowa mountains, Cascade Range, Oregon and Washington.
→ 7
7. Calyces slightly hispid; Cascade Range, Oregon and Washington.
var. flavida
7. Calyces tomentose; Blue and Wallowa mountains, Oregon, and Washington.
var. pachyrhiza
Source FNA vol. 17, p. 521. FNA vol. 17, p. 515.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. bracteosa var. atrosanguinea, P. bracteosa var. bracteosa, P. bracteosa var. canbyi, P. bracteosa var. flavida, P. bracteosa var. latifolia, P. bracteosa var. pachyrhiza, P. bracteosa var. paysoniana, P. bracteosa var. siifolia
Name authority Rydberg: Mem. New York Bot. Gard. 1: 365. (1900) Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 110. (1838)
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