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coil-beak lousewort, coiled lousewort, coiled pedicularis, curve-beak lousewort, curved-beak lousewort, white coiled-beak lousewort

bull elephant's-head, elephant's head, elephant's head pedicularis, elephant's-head lousewort, elephant-head lousewort, pédiculaire du groenland

Habit Plants 10–50 cm. Plants 10–60 cm.
Leaves

basal 3–10, blade lanceolate, 30–100 x 5–30 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, entire or serrate, surfaces glabrous;

cauline 2–10, blade elliptic or triangular to lanceolate, 5–70 x 10–25 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, entire or serrate, surfaces glabrous.

basal 5–20, blade lanceolate, 20–150 x 5–25 mm, 1-pinnatifid or slightly 2-pinnatifid, margins of adjacent lobes nonoverlapping, 1- or 2-serrate, surfaces glabrous;

cauline 3–31, blade lanceolate, 10–150 x 1–25 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous.

Racemes

simple, 1–6, exceeding basal leaves, each 12–40-flowered;

bracts trullate to obtrullate, 5–18 x 2–15 mm, pinnatifid, margins entire, surfaces glabrous.

simple, 1 or 2, exceeding basal leaves, each 20–75-flowered;

bracts linear to trullate, 5–10 x 2–10 mm, undivided to pinnatifid, margins entire, serrate, or 2-serrate, surfaces glabrous.

Pedicels

1.5–5.5 mm.

0.5–1 mm.

Flowers

calyx 5–9 mm, glabrous, lobes 5, triangular, 1–3 mm, apex entire, glabrous;

corolla 7–13 mm, tube white or cream to yellowish or pink to pinkish purple, 4–7 mm;

galea white or cream to yellowish with purple spots at base, or pink to pinkish purple, 2–5.5 mm, beaked, beak coiled, 5–9 mm, base curving, margins entire medially and distally, apex surrounded by abaxial lip, axis of coil nearly vertical;

abaxial lip surrounding beak, white or cream to yellowish, or pink to pinkish purple, 5–8 mm.

calyx 3–5 mm, glabrous or hispid, lobes 5, deltate, 0.5–1.5 mm, apex entire, glabrous;

corolla 5–8 mm, tube purple, rarely white, 3–5 mm;

galea pink to purple, rarely white, 1.5–3 mm, beaked, beak coiled, 5–18 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly horizontal;

abaxial lip pendulous, purple, rarely white, 2–5 mm.

2n

= 16.

Pedicularis contorta

Pedicularis groenlandica

Phenology Flowering Jun–Sep.
Habitat Montane and alpine to arctic bogs, fens, marshes, and forested swamps, seepage areas, stream banks, fens, clay gravel flood plains of rivers.
Elevation 600–3500 m. (2000–11500 ft.)
Distribution
from FNA
CA; ID; MT; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; LB; MB; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The coiled beak that extends directly downward from the top of the galea is distinctive in Pedicularis contorta. Seen from the side, the large, upward-expanded, lateral lobes of the abaxial lip surround the beak, concealing it from view. The flowers of P. contorta are very similar to those of P. racemosa; however, the leaves of P. racemosa are undivided, the beak curves down and to one side, and the inflorescence often branches, forming a panicle of short racemes with long internodes between flowers.

Flower and calyx color and width of the subtending floral bracts delimit three varieties in Pedicularis contorta; corolla color is difficult to discern in herbarium specimens. The following key is adapted from R. N. Reese (1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pedicularis groenlandica has a domed galea, a long, curved beak, and relatively large lateral lobes of the abaxial lip that remarkably resemble the head, trunk, and ears of an elephant. Pedicularis groenlandica is the most widely distributed Pedicularis species in North America; it occurs from the southern Sierra Nevada and southern Rocky Mountains well into the Arctic and Greenland.

Pedicularis attollens and P. groenlandica are sympatric in the Sierra Nevada and the Cascades of central and southern Oregon and can be difficult to distinguish. Pedicularis groenlandica is distinguished by the size and orientation of the beak and color of the corolla; the beak is over twice the length of the beak of P. attollens and is oriented more horizontally compared to the upturned beak of P. attollens. Beak orientation is very difficult to ascertain in herbarium material due to flattening; beak length is a better character on herbarium specimens. The corollas of P. groenlandica are purple with lighter purple to pinkish abaxial lips, whereas the corollas of P. attollens vary from pink to light pink often with conspicuous dark purple spots on the galeas. White-flowered forms of P. groenlandica (forma pallida Lepage) are occasionally seen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corolla tubes, galeas, and abaxial lips white or cream to yellowish, galeas with purple spots at base; calyces pale green.
var. contorta
1. Corolla tubes, galeas, and abaxial lips pink to pinkish purple; calyces green or reddish with purple spots.
→ 2
2. Calyces green; bracts 5–8 mm wide, proximal margins ciliate.
var. ctenophora
2. Calyces reddish with purple spots; bracts 2–4 mm wide, proximal margins glabrous.
var. rubicunda
Source FNA vol. 17, p. 519. FNA vol. 17, p. 523.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. contorta var. contorta, P. contorta var. ctenophora, P. contorta var. rubicunda
Synonyms Elephantella groenlandica, P. groenlandica subsp. surrecta
Name authority Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 108. (1838) Retzius: Fl. Scand. Prodr. ed. 2, 145. (1795)
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