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bract lousewort, fernleaf, towering lousewort, wood-betony

Howell lousewort, Howell's lousewort, Howell's pedicularis

Habit Plants 20–80 cm. Plants 15–40 cm.
Leaves

basal 0–10, blade lanceolate, 20–120 x 10–60 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, 1- or 2-serrate, surfaces glabrous;

cauline 4–10, blade lanceolate, 10–270 x 8–150 mm, undivided or 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate to 2-serrate, surfaces glabrous or scattered glandular.

basal 1–4, blade elliptic to lanceolate, 20–60 x 15–40 mm, undivided or 1-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous;

cauline 10–12, blade ovate to lanceolate, 25–60 x 15–40 mm, undivided or 1-pinnatifid, sometimes auricled, margins of adjacent lobes nonoverlapping or slightly overlapping distally, entire or serrate to crenate, surfaces glabrous or scattered woolly along main vein.

Racemes

simple, 1–4, exceeding basal leaves, each 15–75-flowered;

bracts lanceolate or subulate to trullate, 10–20 x 2–10 mm, undivided, proximal margins entire, distal entire or serrate, surfaces glabrous, hispid, or tomentose.

simple, 1–8, exceeding basal leaves, each 15–40-flowered;

bracts trullate to cordate, 6–8 x 4–8 mm, undivided, proximal margins entire, distal entire, surfaces glabrous or tomentose.

Pedicels

0.5–1 mm.

2.5–3 mm.

Flowers

calyx 7–15 mm, glabrous or tomentose, lobes 5, triangular or filiform, 1–10 mm, apex entire, glabrous or ciliate;

corolla 14–27 mm, tube yellow or dark blood red, 6–12 mm;

galea yellow to yellow tinged with red, purple tinged with yellow, or dark blood red, 6–15 mm, beakless or beaked, beak straight, 0.8–2.5 mm, margins entire medially and distally, apex arching over abaxial lip;

abaxial lip expanded, yellow, yellow tinged with purple, or dark blood red, 4.5–6.5 mm.

calyx 6–6.5 mm, tomentose, lobes 5, triangular, 1.5–2 mm, apex entire, ciliate;

corolla 10–13 mm, tube white, 6–8 mm;

galea white, apically sometimes tinged with red to violet, 3.5–5 mm, beaked, beak straight, 1–2 mm, margins entire medially and distally, apex extending beyond abaxial lip;

abaxial lip white, 1.5–2 mm.

Pedicularis bracteosa

Pedicularis howellii

Phenology Flowering Jun–Aug.
Habitat Alpine forest clearings and edges.
Elevation 1100–2000.
Distribution
from FNA
CA; CO; ID; MT; NM; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 8 (8 in the flora).

Pedicularis bracteosa is found in subalpine habitats across much of western North America, occurring throughout the Rocky Mountains from central British Columbia and Alberta to northern New Mexico, as well as the Coast Range south to northern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pedicularis howellii is found in the Siskiyou Mountains along the California/Oregon border; it has undivided distal leaves. The division of the proximal leaves into irregular and asymmetric lobes, sometimes appearing auricled, is a unique feature of this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. attollens, P. aurantiaca, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. bracteosa var. atrosanguinea, P. bracteosa var. bracteosa, P. bracteosa var. canbyi, P. bracteosa var. flavida, P. bracteosa var. latifolia, P. bracteosa var. pachyrhiza, P. bracteosa var. paysoniana, P. bracteosa var. siifolia
Key
1. Galeas beaked.
→ 2
2. Calyces tomentose.
var. canbyi
2. Calyces glabrous.
var. siifolia
1. Galeas beakless.
→ 3
3. Calyx lobes filiform.
→ 4
4. Corollas: tubes yellow, galeas yellow to yellow tinged with red, or purple tinged with yellow, abaxial lips yellow to yellow tinged with purple.
var. bracteosa
4. Corollas: tubes, galeas, and abaxial lips dark blood red.
var. atrosanguinea
3. Calyx lobes triangular.
→ 5
5. Galeas 10–15 mm.
var. paysoniana
5. Galeas 7–11 mm.
→ 6
6. Galea apices acute; British Colombia, Idaho, Washington.
var. latifolia
6. Galea apices obtuse; Blue and Wallowa mountains, Cascade Range, Oregon and Washington.
→ 7
7. Calyces slightly hispid; Cascade Range, Oregon and Washington.
var. flavida
7. Calyces tomentose; Blue and Wallowa mountains, Oregon, and Washington.
var. pachyrhiza
Name authority Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 110. (1838) A. Gray: Proc. Amer. Acad. Arts 20: 307. (1885)
Source FNA vol. 17, p. 515. Treatment author: Bruce W. Robart. FNA vol. 17, p. 523. Treatment author: Bruce W. Robart.
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