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attol lousewort, attoll lousewort, elephant snouts, little elephant head, little elephant's head, woolly Mammoth

Howell lousewort, Howell's lousewort, Howell's pedicularis

Habit Plants 15–78 cm. Plants 15–40 cm.
Leaves

basal 5–25, blade elliptic, 60–150(or 200–250) x 3–23 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous or scattered glands;

cauline 2–20, blade elliptic, 5–50(–100) x 1–5 mm, undivided or 1(or 2)-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous.

basal 1–4, blade elliptic to lanceolate, 20–60 x 15–40 mm, undivided or 1-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous;

cauline 10–12, blade ovate to lanceolate, 25–60 x 15–40 mm, undivided or 1-pinnatifid, sometimes auricled, margins of adjacent lobes nonoverlapping or slightly overlapping distally, entire or serrate to crenate, surfaces glabrous or scattered woolly along main vein.

Racemes

simple, 1–3, exceeding basal leaves, each 10–50-flowered;

bracts lanceolate to triangular, 5–10 x 3–10 mm, pinnatifid, margins entire, surfaces glabrous or tomentose.

simple, 1–8, exceeding basal leaves, each 15–40-flowered;

bracts trullate to cordate, 6–8 x 4–8 mm, undivided, proximal margins entire, distal entire, surfaces glabrous or tomentose.

Pedicels

1.2–1.6 mm.

2.5–3 mm.

Flowers

calyx 4–5 mm, glabrous or tomentose, lobes 5, triangular, 2–2.5 mm, apex entire, glabrous;

corolla 6–8 mm, tube pink, rarely white, 3–6 mm;

galea white or pink with 2 purple spots or stripes, 1–2 mm, beaked, beak coiled, 3–6 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly vertical;

abaxial lip pendulous, white or pink with purple stripe, 4–5.5 mm.

calyx 6–6.5 mm, tomentose, lobes 5, triangular, 1.5–2 mm, apex entire, ciliate;

corolla 10–13 mm, tube white, 6–8 mm;

galea white, apically sometimes tinged with red to violet, 3.5–5 mm, beaked, beak straight, 1–2 mm, margins entire medially and distally, apex extending beyond abaxial lip;

abaxial lip white, 1.5–2 mm.

2n

= 16.

Pedicularis attollens

Pedicularis howellii

Phenology Flowering Jun–Aug.
Habitat Alpine forest clearings and edges.
Elevation 1100–2000.
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

The flowers of Pedicularis attollens, like those of P. groenlandica, resemble an elephant’s head, and A. Heller placed them both in Elephantella. The short, upturned beak, in contrast to the long, more horizontal downturned beak of P. groenlandica, is a distinguishing feature of P. attollens. Whereas P. groenlandica occurs across much of western and arctic North America, P. attollens is found primarily in the Cascade Range of central and southern Oregon and the Sierra Nevada of California and Nevada. It is also reported from the Klamath Range to the west and the White and Sweetwater mountains and the Warner Range to the east of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pedicularis howellii is found in the Siskiyou Mountains along the California/Oregon border; it has undivided distal leaves. The division of the proximal leaves into irregular and asymmetric lobes, sometimes appearing auricled, is a unique feature of this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 1-pinnatifid.
subsp. attollens
1. Basal leaves 2-pinnatifid.
subsp. protogyna
Source FNA vol. 17, p. 514. FNA vol. 17, p. 523.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. attollens subsp. attollens, P. attollens subsp. protogyna
Synonyms Elephantella attollens
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 384. (1868) A. Gray: Proc. Amer. Acad. Arts 20: 307. (1885)
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