Paysonia |
Brassicaceae tribe Physarieae |
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Habit | Annuals or, rarely, biennials or perennials; not scapose; pubescent or glabrous, trichomes short-stalked, forked or subdendritic, often mixed with coarse, simple ones. | Annuals, biennials, perennials, or subshrubs; eglandular. | ||||||||||||||||||||||||||||
Stems | erect, ascending, decumbent, or procumbent, unbranched or branched. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate, petiolate, blade margins entire, dentate, or pinnately lobed; cauline sessile, blade (base usually auriculate or sagittate, sometimes amplexicaul), margins dentate to lobed. |
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Cauline leaves | petiolate, sessile, or subsessile; blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate. |
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Trichomes | usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones. |
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Racemes | (several-flowered), often strongly elongated in fruit, (rachis straight). |
ebracteate, often elongated in fruit. |
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Flowers | sepals (deciduous), suberect, ascending, or spreading, oblong, (equal), lateral pair not saccate basally, (margins membranous); petals yellow or white, broadly obovate, claw slightly differentiated from blade, (apex rounded, truncate, retuse, or emarginate); stamens (erect), tetradynamous; filaments dilated or not basally; anthers oblong or sagittate, (not apiculate); nectar glands confluent, subtending bases of median stamens, surrounding lateral stamens. |
actinomorphic; sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally; petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct; filaments unappendaged, not winged; pollen (3 or) 4–11-colpate. |
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Fruiting pedicels | ascending to divaricate or recurved, (straight or slightly curved), slender. |
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Fruits | silicles, subsessile or sessile, globose, subglobose, subpyriform, suborbicular, orbicular, elliptic, cordate, or obovoid, terete, flattened, latiseptate, or angustiseptate; valves (papery or leathery), not veined, glabrous or pubescent, (trichomes branched and/or simple); replum not flattened; septum complete, perforated, or reduced to a rim, (membranous); ovules 4–40 per ovary; style distinct, (persistent, slender); stigma capitate, entire. |
silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate; ovules 2–100 per ovary; style usually distinct; stigma entire or strongly 2-lobed. |
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Seeds | sub-biseriate, flattened, margined, orbicular, suborbicular, or oval; cotyledons accumbent. |
biseriate, uniseriate, or aseriate; cotyledons accumbent or incumbent. |
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x | = 7, 8, 9. |
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Paysonia |
Brassicaceae tribe Physarieae |
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Distribution | sc United States; se United States; n Mexico |
North America; Mexico; South America; Asia (ne Russia) |
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Discussion | Species 8 (8 in the flora). Paysonia was segregated from Lesquerella (now Physaria) based on its different base chromosome number, presence of auriculate stem leaves, typically annual duration, trichomes that are neither stellate nor tuberculate, and phylogenetic analysis of DNA sequences. Some species of the genus have very limited distributions and are of conservation concern. Five (P. densipila, P. lescurii, P. lyrata, P. perforata, and P. stonensis) are known to be interfertile under experimental conditions, and some combinations form fertile hybrids in the field (R. C. Rollins 1988). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 7, species ca. 130 (7 genera, 105 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 611. | FNA vol. 7, p. 604. | ||||||||||||||||||||||||||||
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Name authority | O’Kane & Al-Shehbaz: Novon 12: 380. (2002) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895) | ||||||||||||||||||||||||||||
Web links |