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Rugel's nailwort, sand squares

Appalachian nailwort, Virginia nailwort, yellow nailwort

Habit Plants annual; taproot filiform to slender. Plants perennial; caudex branched, woody.
Stems

erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout.

procumbent, branched from base, hirtellose; flowering stems 7–45 cm;

sterile stems 3–10 cm.

Leaves

stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire;

blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent.

stipules narrowly lanceolate, 6–13 mm, apex acuminate, often deeply cleft;

blade linear, 10–30 × 0.4–1 mm, leathery, apex short-spinose, minutely hirtellous to puberulent.

Flowers

5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally;

sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous;

staminodes filiform, widening proximally, 0.5–0.6 mm;

style 1, cleft in distal 1/10.

5-merous, narrowly ovoid, with enlarged hypanthium and calyx tapering gradually distally, 2.8–5.1 mm, glabrous to puberulent, especially proximally;

sepals brown to yellowish, midrib and lateral pair of veins prominent, lanceolate to oblong-lanceolate, 2–2.9 mm, leathery to rigid, margins whitish, 0.1–0.2 mm wide, papery, apex terminated by awn, hood rounded-triangular, awn curved outward, green to red-brown, ± conic, 0.4–1.1 mm, scabrous, distinct spine absent;

staminodes filiform, 0.8–1 mm;

style 1, cleft in distal 1/5, 1.2–2 mm.

Cymes

terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide.

terminal, 3–10+-flowered, somewhat open to compact, often forming clusters 6–30 mm wide.

Utricles

broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous.

ovoid to obovoid, 1.8–2 mm, smooth, glabrous.

Paronychia rugelii

Paronychia virginica

Phenology Flowering late spring–fall. Flowering summer–fall.
Habitat Woodlands, fields, clearings, roadsides, waste places On or among rocks
Elevation 0-200 m (0-700 ft) 700-1300 m (2300-4300 ft)
Distribution
from FNA
FL; GA
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[BONAP county map]
from FNA
AL; AR; DC; MD; MO; NC; OK; TN; TX; VA; WV
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[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cory established var. scoparia for the western populations to reflect the disjunct distribution. Paronychia dichotoma (Linnaeus) Nuttall (1818), sometimes applied to this species, is a later homonym of P. dichotoma de Candolle (1805); see E. L. Core (1940).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 41. FNA vol. 5, p. 42.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. wilkinsonii
Synonyms Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior P. parksii, P. scoparia, P. virginica var. parksii, P. virginica var. scoparia
Name authority (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897) Sprengel: Syst. Veg. 1: 822. (1824)
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