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Rugel's nailwort, sand squares

bristle nailwort

Habit Plants annual; taproot filiform to slender. Plants annual, biennial, or perennial; taproot filiform to slender.
Stems

erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout.

erect to ascending, much-branched distally, 5–20 cm, finely pubescent to puberulent.

Leaves

stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire;

blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent.

stipules lanceolate, 3–8 mm, apex acuminate, entire;

blade linear, 5–20 × 0.3–1 mm, leathery, apex minutely cuspidate, glabrous.

Flowers

5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally;

sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous;

staminodes filiform, widening proximally, 0.5–0.6 mm;

style 1, cleft in distal 1/10.

5-merous, ± short-campanulate, with enlarged hypanthium and calyx widening distally, 2.4–3.2 mm, moderately pubescent proximately with antrorsely appressed to spreading hairs;

sepals green to tan, midrib and lateral pair of veins absent to evident, oblong, 1.2–1.3 mm, leathery to rigid, margins whitish to translucent, 0.05–0.1 mm wide, scarious, apex terminated by awn, hood ± obscure, narrowly rounded, awn widely divergent, 0.6–1.3 mm, broadly conic in proximal 1/3 with white to yellowish, scabrous spine;

staminodes subulate, 0.6–0.7 mm;

style 1, cleft in distal 1/2, 0.4–0.6 mm.

Cymes

terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide.

terminal and lateral, 5–20+-flowered, dichasial and rather diffuse, forming clusters 3–15 mm wide.

Utricles

broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous.

ovoid-oblong, 1 mm, smooth, glabrous.

2n

= 64.

Paronychia rugelii

Paronychia setacea

Phenology Flowering late spring–fall. Flowering spring–fall.
Habitat Woodlands, fields, clearings, roadsides, waste places Limestone barrens, gravelly or sandy slopes and grasslands
Elevation 0-200 m (0-700 ft) 30-200 m (100-700 ft)
Distribution
from FNA
FL; GA
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

We could not distinguish Paronychia lundellorum from P. setacea based on features that Turner used for differentiation, nor did we discover any other discriminating characters. The former was described as perennial (versus annual for the latter) and as having “decided pedicellate” (versus sessile) and longer flowers. Recent data (W. Carr, pers. comm.) indicate that it does bloom in the first year. Furthermore, we have observed that the pedicel character is inconsistent and the flower lengths overlap extensively. Further study of the allopatric populations named P. lundellorum is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 41. FNA vol. 5, p. 42.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. virginica, P. wilkinsonii
Synonyms Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior P. lundellorum, P. setacea var. longibracteata
Name authority (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897) Torrey & A. Gray: Fl. N. Amer. 1: 170. (1838)
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