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Rugel's nailwort, sand squares

fork-chickweed, smooth fork whitlow-wort, smooth-fork nailwort

Habit Plants annual; taproot filiform to slender. Plants annual; taproot filiform to slender.
Stems

erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout.

erect, dichotomously branched, 3–40 cm, ultimate branches filiform, glabrous.

Leaves

stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire;

blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent.

stipules subulate to lanceolate, 0.5–4 mm, apex acuminate, entire;

blade usually dotted or blotched, elliptic to obovate, 3–30 × 1–11 mm, herbaceous, apex acute to obtuse or cuspidate, glabrous.

Flowers

5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally;

sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous;

staminodes filiform, widening proximally, 0.5–0.6 mm;

style 1, cleft in distal 1/10.

5-merous, ± short-cylindric, with enlarged hypanthium and calyx cylindric, 0.9–1.3 mm, glabrous;

sepals greenish to brownish, midrib and lateral pair of veins prominent, ovate to oblong, 0.5–1 mm, herbaceous, margins white to translucent, 0.03–0.1 mm wide, scarious, apex terminated by mucro, hood rounded-triangular, mucro triangular, 0.07–0.15 mm, glabrous;

staminodes absent or indistinct;

styles 2, 0.2–0.3 mm.

Cymes

terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide.

diffuse, flowers solitary (paired) in axils of leafy bracts.

Utricles

broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous.

ovoid to globose, ca. 0.5 mm, granular in distal 1/2, glabrous.

Paronychia rugelii

Paronychia canadensis

Phenology Flowering late spring–fall. Flowering spring–fall.
Habitat Woodlands, fields, clearings, roadsides, waste places Woodlands, thickets, fields, clearings, roadsides, waste places
Elevation 0-200 m (0-700 ft) 0-1300 m (0-4300 ft)
Distribution
from FNA
FL; GA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 41. FNA vol. 5, p. 34.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Synonyms Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior Queria canadensis, Anychia canadensis
Name authority (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897) (Linnaeus) Alph. Wood: Class-Book Bot. ed. s.n.(b), 262. (1861)
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