Paronychia rugelii |
Caryophyllaceae subfam. paronychioideae |
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Rugel's nailwort, sand squares |
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Habit | Plants annual; taproot filiform to slender. | Herbs [small shrubs], annual, biennial, or perennial; taprooted, not rhizomatous. |
Stems | erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout. |
prostrate to ascending or erect, simple or branched. |
Leaves | stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire; blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent. |
opposite, distalmost or all sometimes alternate, bases connate or not, sometimes petiolate, stipulate; stipules ovate or deltate to lanceolate or spatulate, scarious; blade needlelike or often spatulate to elliptic or suborbiculate, seldom succulent. |
Inflorescences | terminal or axillary cymes or flowers solitary; bracts foliaceous or usually scarious; involucel bracteoles absent. |
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Pedicels | present or flowers sessile. |
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Flowers | 5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally; sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous; staminodes filiform, widening proximally, 0.5–0.6 mm; style 1, cleft in distal 1/10. |
bisexual or sometimes unisexual (the plant then dioecious or polygamodioecious); perianth and androecium perigynous; hypanthium usually cup-shaped or cylindric or conic to urceolate; sepals (3–)5, distinct or rarely connate proximally, apex often hooded or awned (awn often subapical); petals absent; stamens absent or 1–5, in 1 whorl arising from hypanthium rim; staminodes absent or 5 (16–19 in Achyronychia); ovary 1-locular; styles 1–3, distinct or sometimes connate proximally; stigmas 2 or 3. |
Fruits | utricles, indehiscent or sometimes opening by 3 or 8–10 valves; carpophore absent. |
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Seeds | 1, white to tan or brown to black, ovoid to reniform, not or slightly laterally compressed; embryo peripheral or central, curved or straight. |
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Cymes | terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide. |
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Utricles | broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous. |
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x | = 7, 8, 9. |
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Paronychia rugelii |
Caryophyllaceae subfam. paronychioideae |
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Phenology | Flowering late spring–fall. | |
Habitat | Woodlands, fields, clearings, roadsides, waste places | |
Elevation | 0-200 m (0-700 ft) | |
Distribution |
FL; GA
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s North America; South America (Andean region); Europe (Mediterranean region); Asia (Mediterranean region, e to India); Africa (Mediterranean region) |
Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 17, species ca. 200 (6 genera, 32 species in the flora). Paronychioideae is characterized by the presence of stipules, petaloid staminodes, and usually indehiscent utricles. It is of similar size to Polycarpoideae; about two-thirds of the species are found in Paronychia and Herniaria. Paronychioideae is sometimes segregated from Caryophyllaceae as Illecebraceae, due to emphasis on the utricle; molecular data does not support recognition of Illecebraceae (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002). While there are some features shared with Polycarpoideae (stipules, solanad type of embryogeny), floral reduction is more pronounced in this group. Tentatively, Corrigioleae (Telephium and Corrigiola) is included here. M. G. Gilbert (1987) proposed transferring this tribe to Molluginaceae, noting that the morphological anomalies within Caryophyllaceae, including alternate leaves, exhibited in these plants were reduced under such an alignment. M. Nepokroeff et al. (2002) retained the tribe within Caryophyllaceae, placed as a sister group to the rest of the family. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 41. | FNA vol. 5, p. 29. |
Parent taxa | Caryophyllaceae > subfam. Paronychioideae > Paronychia | Caryophyllaceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior | |
Name authority | (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897) | Meisner: Pl. Vasc. Gen. 1: 132 |
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