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Rocky Mountain nailwort

Habit Plants perennial, densely cespitose, cushion-forming; caudex much-branched, woody. Herbs [small shrubs], annual, biennial, or perennial; taprooted, not rhizomatous.
Stems

prostrate, much-branched, 5–10 cm, puberulent.

prostrate to ascending or erect, simple or branched.

Leaves

stipules ovate, 3–6 mm, apex subobtuse, entire;

blade narrowly elliptic-oblong or oblong to narrowly elliptic-oblanceolate, 2–5 × 0.2–1.8(–2) mm, fleshy, apex obtuse to subacute, glabrous.

opposite, distalmost or all sometimes alternate, bases connate or not, sometimes petiolate, stipulate;

stipules ovate or deltate to lanceolate or spatulate, scarious;

blade needlelike or often spatulate to elliptic or suborbiculate, seldom succulent.

Inflorescences

terminal or axillary cymes or flowers solitary;

bracts foliaceous or usually scarious;

involucel bracteoles absent.

Pedicels

present or flowers sessile.

Flowers

mostly solitary at end of shoots, almost concealed by leaves.;

flowers 5-merous, elliptic-oblong, with enlarged hypanthium and calyx straight to tapering distally, 2.5–2.8 mm, densely appressed-puberulent, sparsely so distally;

sepals whitish to green, veins absent, narrowly oblong to ovate-oblong, 1.5–1.7 mm, papery to herbaceous, margins white, 0.2–0.3 mm wide, papery, apex with subterminal awn, hood ascending as continuation of sepal, broadly rounded to notched, awn erect, white, 0.3–0.6(–1) mm, ± glabrous spine;

staminodes filiform, 0.7–0.9 mm;

style 1, cleft in distal 1/3, 0.8–1 mm.

bisexual or sometimes unisexual (the plant then dioecious or polygamodioecious);

perianth and androecium perigynous;

hypanthium usually cup-shaped or cylindric or conic to urceolate;

sepals (3–)5, distinct or rarely connate proximally, apex often hooded or awned (awn often subapical);

petals absent;

stamens absent or 1–5, in 1 whorl arising from hypanthium rim;

staminodes absent or 5 (16–19 in Achyronychia);

ovary 1-locular;

styles 1–3, distinct or sometimes connate proximally;

stigmas 2 or 3.

Fruits

utricles, indehiscent or sometimes opening by 3 or 8–10 valves;

carpophore absent.

Seeds

1, white to tan or brown to black, ovoid to reniform, not or slightly laterally compressed;

embryo peripheral or central, curved or straight.

Utricles

ovoid, 1.3–1.5 mm, smooth, glabrous.

x

= 7, 8, 9.

2n

= 32.

Paronychia pulvinata

Caryophyllaceae subfam. paronychioideae

Phenology Flowering summer.
Habitat Rocky slopes and summit screes in alpine regions
Elevation 3300-4200 m (10800-13800 ft)
Distribution
from FNA
CO; UT; WY
[WildflowerSearch map]
[BONAP county map]
s North America; South America (Andean region); Europe (Mediterranean region); Asia (Mediterranean region, e to India); Africa (Mediterranean region)
Discussion

Genera 17, species ca. 200 (6 genera, 32 species in the flora).

Paronychioideae is characterized by the presence of stipules, petaloid staminodes, and usually indehiscent utricles. It is of similar size to Polycarpoideae; about two-thirds of the species are found in Paronychia and Herniaria. Paronychioideae is sometimes segregated from Caryophyllaceae as Illecebraceae, due to emphasis on the utricle; molecular data does not support recognition of Illecebraceae (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002). While there are some features shared with Polycarpoideae (stipules, solanad type of embryogeny), floral reduction is more pronounced in this group.

Tentatively, Corrigioleae (Telephium and Corrigiola) is included here. M. G. Gilbert (1987) proposed transferring this tribe to Molluginaceae, noting that the morphological anomalies within Caryophyllaceae, including alternate leaves, exhibited in these plants were reduced under such an alignment. M. Nepokroeff et al. (2002) retained the tribe within Caryophyllaceae, placed as a sister group to the rest of the family.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 40. FNA vol. 5, p. 29. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Subordinate taxa
Synonyms P. pulvinata var. longiaristata
Name authority A. Gray: Proc. Acad. Nat. Sci. Philadelphia 15: 58. (1864) Meisner: Pl. Vasc. Gen. 1: 132
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