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James' nailwort, nailwort

Rugel's nailwort, sand squares

Habit Plants perennial; caudex branched, woody. Plants annual; taproot filiform to slender.
Stems

erect to ascending, much-branched, 10–35 cm, scabrous, puberulent to pubescent, sometimes glabrous with age.

erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout.

Leaves

stipules lanceolate, 5–15 mm, apex acuminate, entire;

blade linear, 7–25(–34) × 0.5–1 mm, leathery, apex obtuse to subacute or submucronate, pubescent to puberulent.

stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire;

blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent.

Flowers

5-merous, short-campanulate, with enlarged hypanthium and calyx widening somewhat distally, 1.8–2.8 mm, puberulent, glabrous to hirtellous distally;

sepals green to red-brown, veins absent, oblong, 1.3–1.8 mm, leathery to rigid, margins whitish to translucent, 0.05–0.1 mm wide, scarious, apex terminated by awn, broadly rounded, awn widely divergent, 0.4–0.8 mm, conic in proximal 1/2–2/3 with yellowish, scabrous spine;

staminodes filiform, 0.6–1 mm;

style 1, cleft in distal 1/3–1/6, 0.8–1.2 mm.

5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally;

sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous;

staminodes filiform, widening proximally, 0.5–0.6 mm;

style 1, cleft in distal 1/10.

Cymes

terminal, 20–70-flowered, open, clusters 1–2 cm wide.

terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide.

Utricles

ellipsoid-ovoid, 0.8–1 mm, smooth, glabrous.

broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous.

Paronychia jamesii

Paronychia rugelii

Phenology Flowering summer–fall. Flowering late spring–fall.
Habitat Limestone rocky ledges, slopes, hilltops, grasslands Woodlands, fields, clearings, roadsides, waste places
Elevation 500-2500 m (1600-8200 ft) 0-200 m (0-700 ft)
Distribution
from FNA
CO; KS; NE; NM; OK; TX; WY; Mexico (Chihuahua, Coahuila)
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; GA
[WildflowerSearch map]
[BONAP county map]
Discussion

We agree with B. L. Turner (1983b) in not adopting the four varieties of Paronychia jamesii that Chaudhri recognized.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 39. FNA vol. 5, p. 41.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Synonyms P. jamesii var. hirsuta, P. jamesii var. parviflora, P. jamesii var. praelongifolia Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 170. (1838) (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897)
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