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California whitlow wort, San Francisco nailwort, San Francisco whitlow wort

clusterstem nailwort, fork-chickweed, hairy fork nailwort, hairy fork whitlow-wort

Habit Plants perennial, mat-forming; caudex branched, woody. Plants annual; taproot filiform to slender.
Stems

prostrate, usually much-branched throughout, 5–50 cm, pubescent.

erect, usually much-branched, 4–30 cm, retrorsely to spreading-pubescent mostly on 1 side.

Leaves

stipules ovate-lanceolate, 3–6 mm, apex narrowly acute to long-acuminate, entire;

blade elliptic to oblanceolate, 5–10 × 1.5–2.5 mm, ± fleshy, apex spinulous, entire, ± moderately antrorsely appressed-pubescent.

stipules subulate to lanceolate, 0.5–4.5 mm, apex acuminate, sometimes fimbriate;

blade usually dotted or blotched, oblanceolate to elliptic or obovate, 2–25 × 0.5–7(–10) mm, herbaceous to leathery, apex subobtuse to acute or cuspidate, glabrous or occasionally with very few scattered trichomes along midrib.

Flowers

5-merous, short-cylindric, with enlarged hypanthium and calyx cylindric to slightly tapering distally, 1.9–2.4 mm, glabrous, sepals puberulent distally;

sepals greenish, becoming reddish brown, midrib and lateral pair of veins often apparent, oblong to ovate, 1.2–1.3 mm, herbaceous, margins translucent, ca. 0.1 mm wide, scarious, apex terminated by awn, hood broadly rounded, awn erect, 0.5–0.7 mm, conic in proximal 1/6 with whitish, smooth spine;

staminodes absent;

styles 2, 0.2–0.3 mm.

5-merous, short-cylindric, with enlarged hypanthium and calyx ± cylindric, 1.1–1.6 mm, glabrous or with few scattered hairs;

sepals greenish to brownish, veins absent, linear-oblong, 1–1.2 mm, leathery to rigid, margins white to translucent, 0.05–0.1 mm wide, scarious, apex terminated by mucro, hood narrowly rounded, mucro short-conic, 0.05–0.3 mm, ± minutely scabrous;

staminodes absent;

styles 2, 0.07–0.6(–0.7) mm.

Cymes

axillary, inconspicuous, 2–6-flowered, tightly congested.

terminal, 25–70+-flowered, branched, loose to compact, forming clusters 3–10 mm wide.

Utricles

± globose to 4-angled, 1.2–1.3 mm, papillate distally.

obovoid to obconic, 0.7–1 mm, minutely papillate.

2n

= 32, 36.

Paronychia franciscana

Paronychia fastigiata

Phenology Flowering spring.
Habitat Grassy hills
Elevation 20-300 m (100-1000 ft)
Distribution
from FNA
CA; South America (Chile) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Paronychia franciscana was described from California, where it has been known from the San Francisco area since 1887, the species is native in Chile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

The varieties here recognized are sympatric over at least portions of their ranges. A detailed study on infraspecific variation within Paronychia fastigiata is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 0.3-0.6(-0.7) mm, often bent or contorted
var. pumila
1. Styles 0.07-0.25(-0.3) mm, straight
→ 2
2. Sepal cusps 0.2-0.3 mm
var. nuttallii
2. Sepal cusps 0.05-0.15 mm
var. fastigiata
Source FNA vol. 5, p. 38. FNA vol. 5, p. 37.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Subordinate taxa
P. fastigiata var. fastigiata, P. fastigiata var. nuttallii, P. fastigiata var. pumila
Synonyms Anychia fastigiata
Name authority Eastwood: Bull. Torrey Bot. Club 28: 288. (1901) (Rafinesque) Fernald: Rhodora 38: 421. (1936)
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