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California whitlow wort, San Francisco nailwort, San Francisco whitlow wort

paper nailwort, papery whitlow-wort

Habit Plants perennial, mat-forming; caudex branched, woody. Plants annual or short-lived perennial, often densely matted; taproot filiform to slender.
Stems

prostrate, usually much-branched throughout, 5–50 cm, pubescent.

prostrate to ascending, dichotomously branched, 5–20 cm, often retrorsely pubescent on 1 side.

Leaves

stipules ovate-lanceolate, 3–6 mm, apex narrowly acute to long-acuminate, entire;

blade elliptic to oblanceolate, 5–10 × 1.5–2.5 mm, ± fleshy, apex spinulous, entire, ± moderately antrorsely appressed-pubescent.

stipules ovate-lanceolate, 1–1.5 mm, apex obtuse to acuminate, fimbriate;

blade oblong to triangular-ovate, 1.5–6 × 0.5–1.5 mm, leathery, apex acute to obtuse, glabrous.

Flowers

5-merous, short-cylindric, with enlarged hypanthium and calyx cylindric to slightly tapering distally, 1.9–2.4 mm, glabrous, sepals puberulent distally;

sepals greenish, becoming reddish brown, midrib and lateral pair of veins often apparent, oblong to ovate, 1.2–1.3 mm, herbaceous, margins translucent, ca. 0.1 mm wide, scarious, apex terminated by awn, hood broadly rounded, awn erect, 0.5–0.7 mm, conic in proximal 1/6 with whitish, smooth spine;

staminodes absent;

styles 2, 0.2–0.3 mm.

5-merous, occasionally 3- or 4-merous, obconic to short-campanulate, with gradually expanded hypanthium and calyx widening distally, 0.5–0.8 mm, pubescent proximally with straight to hooked hairs;

sepals brown-purple, white to greenish at apex, sometimes proximally, veins absent, ± oblong, 0.5–0.7 mm, leathery to rigid, margins whitish to translucent, 0.1–0.2 mm wide, scarious, apex terminated by crest or mucro, hood broadly rounded, mucro rounded-triangular, ca. 0.1 mm, glabrous;

staminodes absent or rudimentary;

styles 2(–3), ca. 0.3 mm.

Cymes

axillary, inconspicuous, 2–6-flowered, tightly congested.

terminal, 7–25+-flowered, much-branched and very profusely developed along all branches, 1.5–20 mm wide.

Utricles

± globose to 4-angled, 1.2–1.3 mm, papillate distally.

ovoid or ellipsoid to subglobose, 0.4–0.5 mm, smooth, glabrous.

Paronychia franciscana

Paronychia chartacea

Phenology Flowering spring.
Habitat Grassy hills
Elevation 20-300 m (100-1000 ft)
Distribution
from FNA
CA; South America (Chile) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
FL
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Paronychia franciscana was described from California, where it has been known from the San Francisco area since 1887, the species is native in Chile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The occasional occurrence of 3- or 4-merous flowers is unique to Paronychia chartacea among North American species of the genus.

Paronychia chartacea is in the Center for Plant Conservation’s National Collection of Endangered Plants.

The following key is adapted from L. C. Anderson (1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudex (stem base) (1-)1.5-3.5(-4.2) mm; leaf blades 0.5-1.5 mm wide; flower clusters 3-20 mm wide; c peninsular Florida
var. chartacea
1. Caudex (stem base) (0.4-)0.7-1(-1.5) mm; leaf blades 1.2-3 mm wide; flower clusters 1.5-4 mm wide; Florida panhandle
var. minima
Source FNA vol. 5, p. 38. FNA vol. 5, p. 34.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Subordinate taxa
P. chartacea var. chartacea, P. chartacea var. minima
Synonyms Nyachia pulvinata
Name authority Eastwood: Bull. Torrey Bot. Club 28: 288. (1901) Fernald: Rhodora 38: 418. (1936)
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