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clusterstem nailwort, fork-chickweed, hairy fork nailwort, hairy fork whitlow-wort

Rugel's nailwort, sand squares

Habit Plants annual; taproot filiform to slender. Plants annual; taproot filiform to slender.
Stems

erect, usually much-branched, 4–30 cm, retrorsely to spreading-pubescent mostly on 1 side.

erect, dichotomously branched, 5–50 cm, retrorsely to spreading- pubescent throughout.

Leaves

stipules subulate to lanceolate, 0.5–4.5 mm, apex acuminate, sometimes fimbriate;

blade usually dotted or blotched, oblanceolate to elliptic or obovate, 2–25 × 0.5–7(–10) mm, herbaceous to leathery, apex subobtuse to acute or cuspidate, glabrous or occasionally with very few scattered trichomes along midrib.

stipules ovate-lanceolate, 2–6 mm, apex acuminate, entire;

blade linear to linear-oblanceolate or linear-oblong, 3–25 × 1–2.3 mm, herbaceous, apex ± acute, moderately antrorsely puberulent.

Flowers

5-merous, short-cylindric, with enlarged hypanthium and calyx ± cylindric, 1.1–1.6 mm, glabrous or with few scattered hairs;

sepals greenish to brownish, veins absent, linear-oblong, 1–1.2 mm, leathery to rigid, margins white to translucent, 0.05–0.1 mm wide, scarious, apex terminated by mucro, hood narrowly rounded, mucro short-conic, 0.05–0.3 mm, ± minutely scabrous;

staminodes absent;

styles 2, 0.07–0.6(–0.7) mm.

5-merous, cylindric to narrowly lanceolate, with enlarged hypanthium and calyx cylindric to tapering distally, 2.3–3.5 mm, pubescent mostly proximal with hooked or usually straight hairs, sometimes with scattered straight hairs distally;

sepals flecked or solid red-brown, white distally, midrib obscure, linear-lanceolate, 1.5–2 mm, papery, margins white, 0.1–0.2 mm wide, papery, apex terminated by mucro, hood obscure, narrowly rounded, mucro short-conic, ca. 0.1 mm, glabrous;

staminodes filiform, widening proximally, 0.5–0.6 mm;

style 1, cleft in distal 1/10.

Cymes

terminal, 25–70+-flowered, branched, loose to compact, forming clusters 3–10 mm wide.

terminal and lateral, 15–50+-flowered, often forming large, congested clusters 5–30 mm wide.

Utricles

obovoid to obconic, 0.7–1 mm, minutely papillate.

broadly ellipsoid to ± globose, 0.5–0.7 mm, smooth, glabrous.

2n

= 32, 36.

Paronychia fastigiata

Paronychia rugelii

Phenology Flowering late spring–fall.
Habitat Woodlands, fields, clearings, roadsides, waste places
Elevation 0-200 m (0-700 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON; QC
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from FNA
FL; GA
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Discussion

Varieties 3 (3 in the flora).

The varieties here recognized are sympatric over at least portions of their ranges. A detailed study on infraspecific variation within Paronychia fastigiata is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 0.3-0.6(-0.7) mm, often bent or contorted
var. pumila
1. Styles 0.07-0.25(-0.3) mm, straight
→ 2
2. Sepal cusps 0.2-0.3 mm
var. nuttallii
2. Sepal cusps 0.05-0.15 mm
var. fastigiata
Source FNA vol. 5, p. 37. FNA vol. 5, p. 41.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Subordinate taxa
P. fastigiata var. fastigiata, P. fastigiata var. nuttallii, P. fastigiata var. pumila
Synonyms Anychia fastigiata Siphonychia rugelii, Gibbesia rugelii, Odontonychia interior, P. rugelii var. interior, Siphonychia interior
Name authority (Rafinesque) Fernald: Rhodora 38: 421. (1936) (Chapman) Shuttleworth ex Chapman: Fl. South. U.S. ed. 3, 397. (1897)
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