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clusterstem nailwort, fork-chickweed, hairy fork nailwort, hairy fork whitlow-wort

James' nailwort, nailwort

Habit Plants annual; taproot filiform to slender. Plants perennial; caudex branched, woody.
Stems

erect, usually much-branched, 4–30 cm, retrorsely to spreading-pubescent mostly on 1 side.

erect to ascending, much-branched, 10–35 cm, scabrous, puberulent to pubescent, sometimes glabrous with age.

Leaves

stipules subulate to lanceolate, 0.5–4.5 mm, apex acuminate, sometimes fimbriate;

blade usually dotted or blotched, oblanceolate to elliptic or obovate, 2–25 × 0.5–7(–10) mm, herbaceous to leathery, apex subobtuse to acute or cuspidate, glabrous or occasionally with very few scattered trichomes along midrib.

stipules lanceolate, 5–15 mm, apex acuminate, entire;

blade linear, 7–25(–34) × 0.5–1 mm, leathery, apex obtuse to subacute or submucronate, pubescent to puberulent.

Flowers

5-merous, short-cylindric, with enlarged hypanthium and calyx ± cylindric, 1.1–1.6 mm, glabrous or with few scattered hairs;

sepals greenish to brownish, veins absent, linear-oblong, 1–1.2 mm, leathery to rigid, margins white to translucent, 0.05–0.1 mm wide, scarious, apex terminated by mucro, hood narrowly rounded, mucro short-conic, 0.05–0.3 mm, ± minutely scabrous;

staminodes absent;

styles 2, 0.07–0.6(–0.7) mm.

5-merous, short-campanulate, with enlarged hypanthium and calyx widening somewhat distally, 1.8–2.8 mm, puberulent, glabrous to hirtellous distally;

sepals green to red-brown, veins absent, oblong, 1.3–1.8 mm, leathery to rigid, margins whitish to translucent, 0.05–0.1 mm wide, scarious, apex terminated by awn, broadly rounded, awn widely divergent, 0.4–0.8 mm, conic in proximal 1/2–2/3 with yellowish, scabrous spine;

staminodes filiform, 0.6–1 mm;

style 1, cleft in distal 1/3–1/6, 0.8–1.2 mm.

Cymes

terminal, 25–70+-flowered, branched, loose to compact, forming clusters 3–10 mm wide.

terminal, 20–70-flowered, open, clusters 1–2 cm wide.

Utricles

obovoid to obconic, 0.7–1 mm, minutely papillate.

ellipsoid-ovoid, 0.8–1 mm, smooth, glabrous.

2n

= 32, 36.

Paronychia fastigiata

Paronychia jamesii

Phenology Flowering summer–fall.
Habitat Limestone rocky ledges, slopes, hilltops, grasslands
Elevation 500-2500 m (1600-8200 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON; QC
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[BONAP county map]
from FNA
CO; KS; NE; NM; OK; TX; WY; Mexico (Chihuahua, Coahuila)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The varieties here recognized are sympatric over at least portions of their ranges. A detailed study on infraspecific variation within Paronychia fastigiata is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

We agree with B. L. Turner (1983b) in not adopting the four varieties of Paronychia jamesii that Chaudhri recognized.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 0.3-0.6(-0.7) mm, often bent or contorted
var. pumila
1. Styles 0.07-0.25(-0.3) mm, straight
→ 2
2. Sepal cusps 0.2-0.3 mm
var. nuttallii
2. Sepal cusps 0.05-0.15 mm
var. fastigiata
Source FNA vol. 5, p. 37. FNA vol. 5, p. 39.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Paronychia Caryophyllaceae > subfam. Paronychioideae > Paronychia
Sibling taxa
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. franciscana, P. herniarioides, P. jamesii, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
P. ahartii, P. americana, P. argyrocoma, P. baldwinii, P. canadensis, P. chartacea, P. congesta, P. depressa, P. drummondii, P. echinulata, P. erecta, P. fastigiata, P. franciscana, P. herniarioides, P. jonesii, P. lindheimeri, P. maccartii, P. monticola, P. patula, P. pulvinata, P. rugelii, P. sessiliflora, P. setacea, P. virginica, P. wilkinsonii
Subordinate taxa
P. fastigiata var. fastigiata, P. fastigiata var. nuttallii, P. fastigiata var. pumila
Synonyms Anychia fastigiata P. jamesii var. hirsuta, P. jamesii var. parviflora, P. jamesii var. praelongifolia
Name authority (Rafinesque) Fernald: Rhodora 38: 421. (1936) Torrey & A. Gray: Fl. N. Amer. 1: 170. (1838)
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