Paronychia fastigiata |
Caryophyllaceae subfam. paronychioideae |
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clusterstem nailwort, fork-chickweed, hairy fork nailwort, hairy fork whitlow-wort |
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Habit | Plants annual; taproot filiform to slender. | Herbs [small shrubs], annual, biennial, or perennial; taprooted, not rhizomatous. | ||||||||
Stems | erect, usually much-branched, 4–30 cm, retrorsely to spreading-pubescent mostly on 1 side. |
prostrate to ascending or erect, simple or branched. |
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Leaves | stipules subulate to lanceolate, 0.5–4.5 mm, apex acuminate, sometimes fimbriate; blade usually dotted or blotched, oblanceolate to elliptic or obovate, 2–25 × 0.5–7(–10) mm, herbaceous to leathery, apex subobtuse to acute or cuspidate, glabrous or occasionally with very few scattered trichomes along midrib. |
opposite, distalmost or all sometimes alternate, bases connate or not, sometimes petiolate, stipulate; stipules ovate or deltate to lanceolate or spatulate, scarious; blade needlelike or often spatulate to elliptic or suborbiculate, seldom succulent. |
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Inflorescences | terminal or axillary cymes or flowers solitary; bracts foliaceous or usually scarious; involucel bracteoles absent. |
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Pedicels | present or flowers sessile. |
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Flowers | 5-merous, short-cylindric, with enlarged hypanthium and calyx ± cylindric, 1.1–1.6 mm, glabrous or with few scattered hairs; sepals greenish to brownish, veins absent, linear-oblong, 1–1.2 mm, leathery to rigid, margins white to translucent, 0.05–0.1 mm wide, scarious, apex terminated by mucro, hood narrowly rounded, mucro short-conic, 0.05–0.3 mm, ± minutely scabrous; staminodes absent; styles 2, 0.07–0.6(–0.7) mm. |
bisexual or sometimes unisexual (the plant then dioecious or polygamodioecious); perianth and androecium perigynous; hypanthium usually cup-shaped or cylindric or conic to urceolate; sepals (3–)5, distinct or rarely connate proximally, apex often hooded or awned (awn often subapical); petals absent; stamens absent or 1–5, in 1 whorl arising from hypanthium rim; staminodes absent or 5 (16–19 in Achyronychia); ovary 1-locular; styles 1–3, distinct or sometimes connate proximally; stigmas 2 or 3. |
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Fruits | utricles, indehiscent or sometimes opening by 3 or 8–10 valves; carpophore absent. |
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Seeds | 1, white to tan or brown to black, ovoid to reniform, not or slightly laterally compressed; embryo peripheral or central, curved or straight. |
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Cymes | terminal, 25–70+-flowered, branched, loose to compact, forming clusters 3–10 mm wide. |
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Utricles | obovoid to obconic, 0.7–1 mm, minutely papillate. |
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x | = 7, 8, 9. |
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2n | = 32, 36. |
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Paronychia fastigiata |
Caryophyllaceae subfam. paronychioideae |
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Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON; QC
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s North America; South America (Andean region); Europe (Mediterranean region); Asia (Mediterranean region, e to India); Africa (Mediterranean region) |
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Discussion | Varieties 3 (3 in the flora). The varieties here recognized are sympatric over at least portions of their ranges. A detailed study on infraspecific variation within Paronychia fastigiata is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 17, species ca. 200 (6 genera, 32 species in the flora). Paronychioideae is characterized by the presence of stipules, petaloid staminodes, and usually indehiscent utricles. It is of similar size to Polycarpoideae; about two-thirds of the species are found in Paronychia and Herniaria. Paronychioideae is sometimes segregated from Caryophyllaceae as Illecebraceae, due to emphasis on the utricle; molecular data does not support recognition of Illecebraceae (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002). While there are some features shared with Polycarpoideae (stipules, solanad type of embryogeny), floral reduction is more pronounced in this group. Tentatively, Corrigioleae (Telephium and Corrigiola) is included here. M. G. Gilbert (1987) proposed transferring this tribe to Molluginaceae, noting that the morphological anomalies within Caryophyllaceae, including alternate leaves, exhibited in these plants were reduced under such an alignment. M. Nepokroeff et al. (2002) retained the tribe within Caryophyllaceae, placed as a sister group to the rest of the family. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 37. | FNA vol. 5, p. 29. | ||||||||
Parent taxa | Caryophyllaceae > subfam. Paronychioideae > Paronychia | Caryophyllaceae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Anychia fastigiata | |||||||||
Name authority | (Rafinesque) Fernald: Rhodora 38: 421. (1936) | Meisner: Pl. Vasc. Gen. 1: 132 | ||||||||
Web links |