panic raide, switch grass, switch panicgrass
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panic grass, witchgrass
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Plants perennial; rhizomatous, rhizomes often loosely interwoven, hard, with closely overlapping scales, sometimes short or forming a knotty crown. |
Plants annual or perennial; their habit variable. |
40-300 cm tall, 3-5 mm thick, solitary or forming dense clumps, erect or decumbent, usually simple; nodes glabrous; internodes hard, glabrous or glaucous, green or purplish. |
2-300 cm, herbaceous, sometimes hard and almost woody, or woody, simple or branched, bases sometimes cormlike; internodes solid, spongy, or hollow. |
longer than the lower internodes, shorter than those above, glabrous or pilose, especially on the throat, margins usually ciliate; ligules 2-6 mm; blades 10-60 cm long, 2-15 mm wide, flat, erect, ascending or spreading, glabrous or pubescent, adaxial surfaces sometimes densely pubescent, particularly basally, bases rounded to slightly narrowed, margins scabrous. |
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cauline, basal, or both, basal leaves not forming a winter rosette; ligules membranous, usually ciliate; blades filiform to ovate, flat to involute, glabrous or pubescent, cross sections with Kranz anatomy and 1 or 2 bundle sheaths or with non-Kranz anatomy; photosynthesis C4 with NAD-me or NADP-me pathways, or, in plants with non-Krantz anatomy, C3. |
10-55 cm long, 4-20 cm wide, exserted, open; primary branches thin, straight, solitary to whorled or fascicled, ascending to spreading, scabrous, usually rebranching once; pedicels 0.5-20 mm, appressed to spreading. |
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terminal on the culms and branches, often also axillary, terminal panicles typically appearing after midsummer; sterile branches and bristles absent; disarticulation usually below the glumes, sometimes at the base of the upper florets, if at the base of the upper florets, then the florets not very plump at maturity. |
2.5-8 mm long, 1.2-2.5 mm wide, narrowly lanceoloid, turgid to slightly laterally compressed, glabrous, acuminate. |
1-8 mm, usually dorsally compressed, sometimes subterete or laterally compressed, unawned. |
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usually unequal, herbaceous, glabrous or pubescent, rarely tuberculate or glandular, apices not or only slightly gaping at maturity; lower glumes minute to almost equaling the spikelets, 1-9-veined, truncate, acute, or acuminate; upper glumes slightly shorter to much longer than the spikelets, 3-13(15)-veined, bases rarely slightly sulcate, apices rounded to attenuate; lower florets sterile or staminate; lower lemmas similar to the upper glumes; lower paleas absent, or shorter than the lower lemmas and hyaline; upper florets bisexual, sessile or stipitate, apices acute, puberulent, or with a tuft of hairs; upper lemmas usually more or less rigid and chartaceous-indurate, usually shiny, glabrous or (rarely) pubescent, usually smooth, sometimes verrucose or transversely rugose, margins involute, usually clasping the paleas, rarely with basal wings or lunate scars, apices obtuse, acute, apiculate, or with small green crests; upper paleas striate, rarely transversely rugose; lodicules 2; anthers usually 3. |
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smooth; pericarp thin; endosperm hard, without lipid, starch grains simple or compound, or both; hila round or oval, x = 9 (usually), sometimes 10, with polyploid and dysploid derivatives. |
glumes 1.8-3.2 mm, 1/2 - 4/5 as long as the spikelets, glabrous, 5-9-veined, acuminate; upper glumes and lower lemmas extending 0.4-3 mm beyond the upper florets, 7-11-veined, strongly gaping at the apices; lower florets staminate; lower paleas 3-3.5 mm, ovate-hastate, lateral lobes folded over the anthers before anthesis; upper florets 2.3-3 mm long, 0.8-1.1 mm wide, narrowly ovoid, smooth, glabrous, shiny; upper lemmas clasping the paleas only at the base. |
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= 18, 21, 25, 30, 32, 35, 36, 54-60, 67-72, 74, 77, 90, 108. |
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AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; WY; HI; MB; NS; ON; QC; SK
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AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; AB; BC; LB; MB; NB; NS; ON; PE; QC; SK; Virgin Islands |
Panicum virgatum grows in tallgrass prairies, especially mesic to wet types where it is a major component of the vegetation, and on dry slopes, sand, open oak or pine woodlands, shores, river banks, and brackish marshes. Its range extends, primarily on the eastern side of the Rocky Mountains, from southern Canada through the United States to Mexico, Cuba, Bermuda, and Costa Rica, and, possibly as an introduction, in Argentina. It has also been introduced as a forage grass to other parts of the world. Panicum virgatum is an important and palatable forage grass, but its abundance in native grasslands decreases with grazing. Several types are planted for range and wildlife habitat improvement. Plants from eastern New Mexico, western Texas, and northern Mexico tend to have larger spikelets (6-8 mm versus 2.5-5.5 mm) and are sometimes called P. havardii Vasey Tetraploids appear to be the most common ploidy level, especially in the upper midwest and northern plains, with higher ploidy levels being more common southwards, but plants in a small area can range from diploid through duodecaploid, with dysploid derivatives. If morphological markers matched chromosome numbers and ecotypic characters, the species could be considered an aggregate of numerous microspecies. In the absence of such correlations, it must be regarded as simply a wide-ranging, highly variable taxon. Plants identified as Panicum virgatum var. cubense Griseb. and P. virgatum var. spissum Linder represent end points of geographic clines. Panicum virgatum is not always readily separable from P. amarum, particularly P. amarum subsp. amarulum; future work may support their treatment as conspecific taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Panicum is a large genus, but just how large is difficult to estimate because its limits are not yet clear. Many taxonomists would treat it as including Dichanthelium, Steincbisma, and some members of Urochloa. Recent work supports some aspects of the treatment presented here, but not all of them. For instance, Guissani et al. (2001) suggest that Panicum subg. Panicum is a monophyletic group that should have a rank equivalent to Dichanthelium and Steinchisma. The two other subgenera included here in Panicum, subg. Agrostoidea and subg. Phanopyrum, are not monophyletic, but the relationships of their species to other members of Panicum sensu lato are not well enough understood to suggest a better treatment, nor to justify the name changes a differing generic treatment would require. Most species of Panicum are tropical, but many grow in warm, temperate regions. Of the thirty-four species occurring in the Flora region, twenty-five are native, seven are established introductions, and two are not established within the region. Within the Flora region, Panicum is most abundant in the southeastern United States. Many species grow in early serai stages or weedy areas; some grow at forest edges, in prairies, savannahs, deserts, forests, beaches, and in shallow water. Panicum miliaceum has been grown since prehistory in China and India as a cereal grain, and is a common component of bird seed. Seeds of P. hirticaule subsp. sonorum have been used for food by the Cocopa tribe of the southwest. Important hay and range species include P. virgatum, P. rigidulum, P. bulbosum, P. obtusum, and P. repens. Apomixis, polyploidy, and autogamy have produced numerous microspecies in some groups; hybridization and introgression has resulted in a reticulum of intergrading forms in some complexes. The number of taxa recognized has varied widely over the past century. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Panicle branches 1-sided; spikelets usually subsessile, the longest pedicels usually less than 2 mm long, rarely 3 mm long. | → 2 |
2. Spikelets 5.5-7 mm long; upper florets less than 1/3 as long as the spikelets (sect. Phanopyrum) | P. gymnocarpon |
2. Spikelets 1.6-4.4 mm long; upper florets 2/5 as long as to almost equaling the spikelets. | → 3 |
3. Lower glumes 5- or 7-veined, about 3/4 as long as the spikelets; plants with stolons or shallow rhizomes (sect. Obtusa) | P. obtusum |
3. Lower glumes 1- or 3-veined, 1/2 - 2/3 as long as the spikelets; plants without stolons, often with rhizomes. | → 4 |
4. Lower florets staminate; lower paleas subequal to the lower lemmas; upper thin, lemmas flexible, clasping the paleas only at the base (sect. Hemitonia) | P. hemitomon |
4. Lower florets sterile; lower paleas no more than 2/3 as long as the lower lemmas; upper lemmas thick, stiff, clasping the paleas throughout their length. | → 5 |
5. Glumes and lower lemmas without keeled midveins; upper florets with glabrous apices; plants tufted, from knotty rhizomes; panicles with a few spikelets; pedicels with slender hairs near the apices (sect. Tenera) | P. tenerum |
5. Glumes and lower lemmas with keeled midveins; upper florets with a tuft of small hairs at the apices; plants often with scaly rhizomes; panicles with many spikelets; pedicels glabrous (sect. Agrostoidea). | → 6 |
6. Plants without conspicuous rhizomes, cespitose; culms and sheaths strongly compressed; spikelets usually 1.6-3.8 mm long, lanceolate, not falcate | P. rigidulum |
6. Plants with conspicuous, stout, short or elongate, scaly rhizomes; culms and sheaths slightly compressed; spikelets 2.3-3.9 mm long, rarely lanceolate, often falcate | P. anceps |
1. Panicle branches usually not 1-sided; spikelets not subsessile, the longest pedicels 2-20 mm long. | → 7 |
7. Upper glumes and lower lemmas warty-tuberculate (sect. Verrucosa). | → 8 |
8. Lower lemmas verrucose with hemispheric warts; spikelets 1.7-2.2 mm long, about 1 mm wide, subacute or obtuse, glabrous; plants of wetlands | P. verrucosum |
8. Lower lemmas tuberculate-hispid; spikelets 3.2-4 mm long, about 1.5 mm wide, acute or acuminate; plants of dry, sandy or clayey areas | P. brachyanthum |
7. Upper glumes and lower lemmas glabrous, villous, or scabridulous, but not warty-tuberculate. | → 9 |
9. Upper florets faintly to evidently transversally rugose; sheaths keeled; culm bases often cormlike (sect. Bulbosa). | → 10 |
10. Culm bases thickened, cormlike; culms slightly compressed; rhizomes, if present, short and thin; spikelets 2.8-5.4 mm long; lower glumes 1.2-3.5 mm long, 1/2 - 4/5 as long as the spikelets | P. bulbosum |
10. Culm bases not cormlike; culms strongly compressed; rhizomes present, long, stout; spikelets 2.5-3.4 mm long; lower glumes usually less than 1.7 mm long, up to 1/2 as long as the spikelets | P. plenum |
9. Upper florets smooth or striate, rarely inconspicuously rugose; sheaths not keeled; culm bases never cormlike. | → 11 |
11. Plants with rhizomes about 1 cm thick and with large, pubescent, scalelike leaves; culms hard, almost woody (sect. Antidotalia) | P. antidotale |
11. Plants without rhizomes or with rhizomes less than 0.5 cm thick and with small, glabrous, scalelike leaves; culms clearly not woody, except at the base of P. hirsutum (subg. Panicum). | → 12 |
12. Glumes, lower lemmas, and upper lemma margins villous, with whitish hairs (sect. Urvilleana) | P. urvilleanum |
12. Glumes and lemmas usually glabrous, sometimes the lower lemmas sparsely pilose on the margins and near the apices. | → 13 |
13. Plants perennial, usually with vigorous scaly rhizomes; lower florets staminate (sect. Repentia). | → 14 |
14. Lower glumes 0.5-1.5 mm long, less than 1/2 as long as the spikelet, 1-5-veined; upper glumes and lower lemmas extending 0.1-0.5 mm beyond the upper florets and scarcely separated (gaping); lower paleas oblong, not hastate-lobed. | → 15 |
15. Lower glumes subtruncate to broadly acute, faintly veined; upper florets widest at or above the middle, with rounded apices; plants not cespitose, with long, scaly rhizomes | P. repens |
15. Lower glumes acute, with evident veins; upper florets widest below the middle, with lightly beaked apices; plants cespitose, with short knotty rhizomes | P. coloratum |
14. Lower glumes 1.8-4 mm long, more than 1/2 as long as the spikelets, with at least 5 veins; upper glumes and lower lemmas extending 0.4-3 mm beyond the upper florets, stiffly separated (gaping); lower paleas hastate-lobed. | → 16 |
16. Panicles contracted; branches appressed to strongly ascending; plants glabrous throughout | P. amarum |
16. Panicles open; branches ascending to spreading; plants often pilose, at least at the base of the leaf blades | P. virgatum |
13. Plants annual, or perennials usually without rhizomes, sometimes rooting at the lower nodes; lower florets sterile. | → 17 |
17. Lower glumes truncate to subacute, 1/5 – 1/3 as long as the spikelets; sheaths more or less compressed, glabrous or sparsely pubescent; plants slightly succulent or spongy (sect. Dichotomiflora). | → 18 |
18. Plants usually annual, usually terrestrial, rooting at the lower nodes if in water, but not floating; blades 3-25 mm wide | P. dichotomiflorum |
18. Plants perennial or of indefinite duration, usually aquatic, sometimes floating, rooting at the lower nodes; blades 2-15 mm wide. | → 19 |
19. Spikelets 2-2.2 mm long; blades 2-4 mm wide; lower paleas absent; culms succulent | P. lacustre |
19. Spikelets 3-4 mm long; blades 5-15 mm wide; lower paleas present; culms spongy | P. paludosum |
17. Lower glumes acute to attenuate, usually 1/3 – 3/4 as long as the spikelets; sheaths rounded, usually hirsute or hispid; plants not succulent (sect. Panicum). | → 20 |
20. Spikelets 4-6.5 mm long. | → 21 |
21. Upper glumes and lower lemmas only slightly exceeding the upper florets; upper florets 2-2.5 mm wide; plants annual; lower paleas truncate to bilobed | P. miliaceum |
21. Upper glumes and lower lemmas exceeding the upper florets by 3-4 mm; upper florets 1-1.1 mm wide; plants perennial; lower paleas acute | P. capillarioides |
20. Spikelets 1-4.2 mm long. | → 22 |
22. Plants perennial; panicle branches usually with all or most secondary branches confined to the distal 1/3. | → 23 |
23. Lower panicle branches whorled; culms 2-10 mm thick, 50-300 cm tall. | → 24 |
24. Sheaths with fragile, prickly hairs causing skin irritation; panicles not breaking at the base and becoming tumble-weeds; lower paleas 1.3-1.7 mm long | P. hirsutum |
24. Sheaths glabrous or sparsely to densely pubescent but without fragile, prickly hairs; panicles breaking at the base and becoming tumbleweeds; lower paleas 1.4-2.2 mm long | P. bergii |
23. Lower panicle branches solitary; culms 0.5-10 mm thick, 15-100 cm tall. | → 25 |
25. Blades glabrous and glaucous on the adaxial surface; nodes sericeous or pilose, sometimes almost glabrous | P. ballii |
25. Blades sparsely to densely hirsute and not glaucous on the adaxial surface; nodes sericeous. | → 26 |
26. Spikelets 2.1-2.9 mm long; culms spreading to weakly ascending; blades spreading, 1-5 mm wide, without a prominent white midrib | P. diffusum |
26. Spikelets 2.6-3.4 mm long; culms erect to decumbent; blades ascending to erect, 0.5-14 mm wide, with a prominent white midrib | P. ghiesbreghtii |
22. Plants annual; panicle branches usually with secondary branches and pedicels attached to the distal 2/3 | → 27 |
27. Blades 2-7 cm long, 5-20 mm wide, lanceolate, 4-6 times longer than wide (sect. Monticola, in part) | P. trichoides |
27. Blades 5-40 cm long, 1-18 mm wide, linear, more than 10 times longer than wide (sect. Panicum, in part). | → 28 |
28. Panicles more than 2 times longer than wide at maturity; branches ascending to somewhat divergent; spikelets narrowly ovoid, usually about 3 times longer than wide | P. flexile |
28. Panicles less than 1.5 times longer than wide at maturity; branches diverging; spikelets variously shaped, less than 3 times longer than wide. | → 29 |
29. Spikelets 2.1-4 mm long, upper glumes and lower lemmas with prominent veins; lower glumes 2/5 – 3/4 as long as the spikelets; lower paleas 0.4-2 mm long, from 1/3 as long as the lower lemmas to equaling them; ligules 0.2-0.4 mm or 1-3.5 mm long. | → 30 |
30. Lower glumes 0.7-1.1 mm long, about 2/5 as long as the spikelets; lower paleas 1-2 mm long; leaf blades 2-8 mm wide, usually completely glabrous, sometimes with a few marginal cilia near the base | P. psilopodium |
30. Lower glumes 1.2-2.4 mm long, 1/2 - 3/4 as long as the spikelets; lower paleas 0.2-0.9 mm long; leaf blades 1-30 mm wide, hairs papillose-based. | → 31 |
31. Primary panicle branches appressed to the main axis; culms 2-8 cm long; spikelets 2-2.2 mm long | P. mobavense |
31. Primary panicle branches divergent; culms 11-110 cm long; spikelets 1.9-4 mm long | P. hirticaule |
29. Spikelets 1.4-4 mm long, upper glumes and lower lemmas without prominent veins; lower glumes usually less than 1/2 as long as the spikelets; lower paleas usually small or absent; ligules 0.5-1.5 mm long. | → 32 |
32. Plants mostly glabrous, but the sheaths ciliate on the margins and the blades sometimes sparingly pilose adaxially (sect. Monticola, in part) | P. bisulcatum |
32. Plants mostly hairy, even the sheaths hairy throughout. | → 33 |
33. Panicles usually more than 1/2 the total height of the plant, breaking at the base of the peduncle at maturity and becoming a tumbleweed; spikelets 1.9-4 mm long; mature upper florets stramineous or nigrescent (sect. Panicum, in part) | P. capillare |
33. Panicles usually less than 1/2 the total height of the plant, the base of the peduncle usually not breaking at maturity; spikelets 1.4-2.4 mm long; mature upper florets often dark brown | P. philadelphicum |
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FNA vol. 25, p. 474. |
FNA vol. 25, p. 450. Author: Robert W. Freckmann; Michel G. Lelong;. |
Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia |
Poaceae > subfam. Panicoideae > tribe Paniceae |
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum |
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P. amarum, P. anceps, P. antidotale, P. ballii, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mobavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum |
P. virgatum var. spissum, P. virgatum var. cubense, P. bavardii |
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L. |
L. |
- Local floras: OR
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| - Local floras: CA, OR, WA
- Local Web sites: CalFlora, CalPhotos, Flora NW, IL Wildflowers, KS Wildflowers, LA Plants, MD Biodiversity, MI Flora, MN Wildflowers, MO Plants, PNW Herbaria
- WildflowerSearch
- iNaturalist (observations)
- USDA Plants Database
- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
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