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panic de philadelphie, Philadelphia panic grass, Philadelphia witchgrass

couch panicum, creeping panic, panic rampant, torpedo grass, wainaku grass

Habit Plants annual; hirsute, hairs papillose-based, usually yellow-green to green, sometimes purplish. Plants perennial; rhizomatous, forming extensive colonies, rhizomes long, to 5 mm thick, branching, scaly, sharply pointed.
Culms

8-100 cm tall, about 1 mm thick, erect to decumbent, simple to profusely branched;

nodes sparsely to densely pilose.

20-90 cm tall, 1.8-2.8 mm thick, erect, rigid, simple or branching from the lower and middle nodes;

nodes glabrous or sparsely hispid;

internodes glabrous.

Sheaths

generally shorter than the internodes, not keeled, lower nodes glabrous or hispid, hairs papillose-based, particularly near the summits;

ligules 0.5-1 mm;

blades 3-25 cm long, 2-8 mm wide, often distichous, flat to slightly involute, firm, adaxial surfaces pilose basally, glabrous or sparsely pubescent abaxially.

Leaves

often crowded basally;

sheaths rounded, usually longer than the internodes, hispid, hairs papillose-based, to 5 mm;

ligules 0.5-1.5 mm;

blades 3-30 cm long, 2-12 mm wide, linear, ascending to erect, flat, hirsute to sparsely pilose, greenish or purplish, bases truncate to subcordate and ciliate on the margins, apices acute.

Panicles

7-27 cm long, 4-24 cm wide, 1/4 - 1/3 as long as the plants, diffuse, usually exserted at anthesis, not breaking at the base of the peduncles to become a tumbleweed;

rachises glabrous or sparsely pilose basally;

primary branches spreading, secondary branches and pedicels confined to the distal 2/3 secondary branches diverging to appressed, with 1-4 spikelets;

pedicels 3-15 mm, spreading to appressed, scabrous or hirsute;

pulvini glabrous or pilose.

3-24 cm long, usually less than 5 cm wide, open;

primary branches 2-11 cm, alternate, few, stiffly ascending to spreading;

pedicels 1-6 mm, subappressed.

Spikelets

1.4-2.4 mm long, 0.5-0.7 mm wide, usually green, glabrous.

2.2-2.8 mm long, 0.8-1.3 mm wide, ellipsoid-ovoid, pale green, acute, upper glumes and lower lemmas sometimes separating (gaping) beyond the florets.

Lower glumes

0.5-0.9 mm, usually less than 1/2 as long as the spikelets, 3-4-veined, truncate to acuminate;

upper glumes 1.6-2 mm, 7-veined, veins not prominent;

lower lemmas 1.6-1.9 mm, 7-9-veined, veins not prominent;

lower paleas absent;

lower florets sterile;

upper florets 1.5-1.7 mm long, about 0.4 mm wide, often dark brown, sometimes disarticulating, apices minutely papillose.

0.5-1 mm, 1/5 – 2/5 as long as the spikelets, glabrous, faintly 1-5-veined, subtruncate to broadly acute;

upper glumes and lower lemmas glabrous, extending 0.1-0.5 mm beyond the upper florets, scarcely separated;

upper glumes 7-11-veined, shorter than the lower lemmas, acute to short-acuminate;

lower florets staminate;

lower lemmas 7-11-veined;

lower paleas 1.9-2.1 mm, oblong;

upper florets 1.8-2.7 mm long, 0.7-1.3 mm wide, broadly ellipsoid, broadest at or above the middle, glabrous, shiny, smooth, apices rounded.

2n

= 18.

- 36, 40, 45, 54.

Panicum philadelphicum

Panicum repens

Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CA; FL; GA; LA; MS; NC; SC; TX; HI
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum philadelphicum grows in open areas such as fallow fields, roadside ditches, receding shores, and rock crevices. It is restricted to the eastern part of the Flora region. It intergrades with P. capillare, possibly as a result of hybridization, especially in the southeastern United States. Seeds germinating on receding shores in late summer often produce tiny plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum repens grows on open, moist, sandy beaches and the shores of lakes and ponds, occasionally extending out into or onto the water. It is mostly, but not exclusively, coastal. It grows on tropical and subtropical coasts throughout the world and may have been introduced to the Americas from elsewhere. Small plants having small, dense panicles of purplish spikelets with longer, subacute lower glumes have been named Panicum gouinii E. Fourn., but they intergrade with more typical plants and do not seem to merit taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets less than 1/2 as wide as long; plants purplish
subsp. lithophilum
1. Spikelets usually more than 1/2 as wide as long; plants green or yellow-green.
→ 2
2. Spikelets 1.9-2.4 mm long; apices of the upper glumes and lower lemmas straight; secondary branches and pedicels divergent; blades often 6-12 mm wide, those of the flag leaves usually more than 1/2 as long as the panicles
subsp. gattingeri
2. Spikelets 1.4-2.1 mm long; apices of the upper glumes and lower lemmas curving over the upper florets at maturity; secondary panicle branches and pedicels appressed; blades usually 2-6 mm wide, those of the flag leaves usually less than M as long as the panicles
subsp. philadelphicum
Source FNA vol. 25, p. 459. FNA vol. 25.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia
Sibling taxa
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. philadelphicum subsp. gattingeri, P. philadelphicum subsp. lithophilum, P. philadelphicum subsp. philadelphicum
Synonyms P. tuckermanii
Name authority Bernh. ex Trin. L.
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