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fall panicgrass, fall panicum, knee grass, panic d'automne, smooth witchgrass

panic de philadelphie, Philadelphia panic grass, Philadelphia witchgrass

Habit Plants annual or short-lived perennials in the Flora region, perennial in the tropics; usually terrestrial, sometimes aquatic but not floating. Plants annual; hirsute, hairs papillose-based, usually yellow-green to green, sometimes purplish.
Culms

5-200 cm tall, 0.4-3 mm thick, decumbent to erect, commonly geniculate to ascending, rooting at the lower nodes when in water, simple to divergently branched from the lower and middle nodes, usually succulent, slightly compressed, glabrous;

nodes usually swollen, sometimes constricted on robust plants, glabrous;

internodes glabrous, shiny, pale green to purplish.

8-100 cm tall, about 1 mm thick, erect to decumbent, simple to profusely branched;

nodes sparsely to densely pilose.

Sheaths

compressed, inflated, sparsely pubescent near the base, elsewhere mostly glabrous, sparsely pilose, or hispid, hairs sometimes papillose-based, margins or throat ciliate, with papillose-based hairs;

ligules 0.5-2 mm;

blades 10-65 cm long, 3-25 mm wide, glabrous or sparsely pilose, often scabrous near the margins, midribs stout, whitish.

Leaves

often crowded basally;

sheaths rounded, usually longer than the internodes, hispid, hairs papillose-based, to 5 mm;

ligules 0.5-1.5 mm;

blades 3-30 cm long, 2-12 mm wide, linear, ascending to erect, flat, hirsute to sparsely pilose, greenish or purplish, bases truncate to subcordate and ciliate on the margins, apices acute.

Panicles

4-40 cm, diffuse, lax, with a few spikelets;

branches to 15 cm, alternate or opposite, occasionally verticillate, ascending to spreading, stiff, scabrous;

pedicels 1-6 mm, sharply 3-angled, scabrous, expanded to cuplike apices, appressed mostly to the abaxial side of the branches.

7-27 cm long, 4-24 cm wide, 1/4 - 1/3 as long as the plants, diffuse, usually exserted at anthesis, not breaking at the base of the peduncles to become a tumbleweed;

rachises glabrous or sparsely pilose basally;

primary branches spreading, secondary branches and pedicels confined to the distal 2/3 secondary branches diverging to appressed, with 1-4 spikelets;

pedicels 3-15 mm, spreading to appressed, scabrous or hirsute;

pulvini glabrous or pilose.

Spikelets

1.8-3.8 mm long, 0.7-1.2 mm wide, ellipsoid to narrowly ovoid, light green to red-purple, glabrous, acute to acuminate.

1.4-2.4 mm long, 0.5-0.7 mm wide, usually green, glabrous.

Lower glumes

0.6-1.2 mm, 1/4 - 1/3 as long as the spikelets, 0-3-veined, obtuse to acute;

upper glumes and lower lemmas similar, exceeding the upper florets by 0.3-0.6 mm, 7-9-veined;

lower paleas vestigial to almost as long as the lower lemmas;

lower florets sterile;

upper florets 1.4-2.5 mm long, 0.7-1.1 mm wide, narrowly ellipsoid, smooth, shiny, stramineous to nigrescent, with pale veins.

0.5-0.9 mm, usually less than 1/2 as long as the spikelets, 3-4-veined, truncate to acuminate;

upper glumes 1.6-2 mm, 7-veined, veins not prominent;

lower lemmas 1.6-1.9 mm, 7-9-veined, veins not prominent;

lower paleas absent;

lower florets sterile;

upper florets 1.5-1.7 mm long, about 0.4 mm wide, often dark brown, sometimes disarticulating, apices minutely papillose.

2n

= 36, 54.

= 18.

Panicum dichotomiflorum

Panicum philadelphicum

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; HI; PR; BC; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum dichotomiflorum grows in open, often wet, disturbed areas such as cultivated and fallow fields, roadsides, ditches, open stream banks, receding shores, clearings in flood plain woods, and sometimes in shallow water. It is probably native throughout the eastern United States and adjacent Canada, but introduced elsewhere, including in the western United States. Its size and habit may be partly under genetic control, but these features also seem to be strongly affected by moisture levels, soil richness, competition, and the time of germination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum philadelphicum grows in open areas such as fallow fields, roadside ditches, receding shores, and rock crevices. It is restricted to the eastern part of the Flora region. It intergrades with P. capillare, possibly as a result of hybridization, especially in the southeastern United States. Seeds germinating on receding shores in late summer often produce tiny plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 1.8-2.2 mm long, widest at the middle, acute; upper glumes and lower lemmas submembranaceous; pedicels often over 3 mm long
subsp. puritanorum
1. Spikelets 2.2-3.8 mm long, widest below the middle, acuminate; upper glumes and lower lemmas subcoriaceous; most pedicels less than 3 mm long.
→ 2
2. Sheaths glabrous or sparsely pilose, hairs not papillose-based
subsp. dichotomiflorum
2. Sheaths hispid, hairs papillose-based
subsp. bartowense
1. Spikelets less than 1/2 as wide as long; plants purplish
subsp. lithophilum
1. Spikelets usually more than 1/2 as wide as long; plants green or yellow-green.
→ 2
2. Spikelets 1.9-2.4 mm long; apices of the upper glumes and lower lemmas straight; secondary branches and pedicels divergent; blades often 6-12 mm wide, those of the flag leaves usually more than 1/2 as long as the panicles
subsp. gattingeri
2. Spikelets 1.4-2.1 mm long; apices of the upper glumes and lower lemmas curving over the upper florets at maturity; secondary panicle branches and pedicels appressed; blades usually 2-6 mm wide, those of the flag leaves usually less than M as long as the panicles
subsp. philadelphicum
Source FNA vol. 25, p. 469. FNA vol. 25, p. 459.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Dichotomiflora Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. dichotomiflorum subsp. bartowense, P. dichotomiflorum subsp. dichotomiflorum, P. dichotomiflorum subsp. puritanorum
P. philadelphicum subsp. gattingeri, P. philadelphicum subsp. lithophilum, P. philadelphicum subsp. philadelphicum
Synonyms P. tuckermanii
Name authority Michx. Bernh. ex Trin.
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