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beak panicgrass

panic de philadelphie, Philadelphia panic grass, Philadelphia witchgrass

Habit Plants perennial; conspicuously rhizomatous, rhizomes short or elongate, stout, scaly. Plants annual; hirsute, hairs papillose-based, usually yellow-green to green, sometimes purplish.
Culms

30-130 cm, terete to slightly compressed.

8-100 cm tall, about 1 mm thick, erect to decumbent, simple to profusely branched;

nodes sparsely to densely pilose.

Sheaths

laterally compressed, glabrous or sparsely to densely pilose or villous, especially at the summit;

ligules less than 0.5 mm, membranous, erose, often brownish;

blades 10-50 cm long, 4-12 mm wide, erect, adaxial surfaces pilose at least basally, glabrous or pilose abaxially.

Leaves

often crowded basally;

sheaths rounded, usually longer than the internodes, hispid, hairs papillose-based, to 5 mm;

ligules 0.5-1.5 mm;

blades 3-30 cm long, 2-12 mm wide, linear, ascending to erect, flat, hirsute to sparsely pilose, greenish or purplish, bases truncate to subcordate and ciliate on the margins, apices acute.

Panicles

10-40 cm, 1/4 - 2/3 as wide as long, well-exserted at anthesis;

branches relatively few, stiffly spreading or ascending; ultimate branchlets 1-sided;

pedicels 0.1-3 mm, scabridulous to scabrous, appressed.

7-27 cm long, 4-24 cm wide, 1/4 - 1/3 as long as the plants, diffuse, usually exserted at anthesis, not breaking at the base of the peduncles to become a tumbleweed;

rachises glabrous or sparsely pilose basally;

primary branches spreading, secondary branches and pedicels confined to the distal 2/3 secondary branches diverging to appressed, with 1-4 spikelets;

pedicels 3-15 mm, spreading to appressed, scabrous or hirsute;

pulvini glabrous or pilose.

Spikelets

2.3-3.9 mm, narrowly ellipsoid to ovoid, usually subsessile, usually pale to yellowish-green, glabrous, often falcate and gaping at the apices, rarely lanceolate, densely crowded on short, appressed branchlets, set obliquely on short pedicels.

1.4-2.4 mm long, 0.5-0.7 mm wide, usually green, glabrous.

Lower glumes

A-A as long as the spikelets, 3-veined, keels scabrous, apices acute;

upper glumes and lower lemmas subequal, keeled, beaked, usually gaping at the apices;

lower florets sterile;

lower paleas subequal to the lower lemmas;

upper florets 1.5-2.2 mm long, about 1 mm wide, 2/5 – 3/4 as long as the spikelets, apices with a tuft of minute, thick hairs;

upper lemmas thick, stiff, clasping the upper paleas throughout their length.

0.5-0.9 mm, usually less than 1/2 as long as the spikelets, 3-4-veined, truncate to acuminate;

upper glumes 1.6-2 mm, 7-veined, veins not prominent;

lower lemmas 1.6-1.9 mm, 7-9-veined, veins not prominent;

lower paleas absent;

lower florets sterile;

upper florets 1.5-1.7 mm long, about 0.4 mm wide, often dark brown, sometimes disarticulating, apices minutely papillose.

2n

= 18, 36.

= 18.

Panicum anceps

Panicum philadelphicum

Distribution
from FNA
AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV
[WildflowerSearch map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum anceps grows in low, moist, primarily sandy areas, pine savannahs, the borders of flood-plain swamps, mesic woodlands, roadsides, and upland pine-hardwood forests. It is restricted to the United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum philadelphicum grows in open areas such as fallow fields, roadside ditches, receding shores, and rock crevices. It is restricted to the eastern part of the Flora region. It intergrades with P. capillare, possibly as a result of hybridization, especially in the southeastern United States. Seeds germinating on receding shores in late summer often produce tiny plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 2.7-3.9 mm long, often clearly falcate; rhizomes relatively short and stout
subsp. anceps
1. Spikelets 2.3-2.8 mm long, not clearly falcate; rhizomes relatively long and slender
subsp. rhizomatum
1. Spikelets less than 1/2 as wide as long; plants purplish
subsp. lithophilum
1. Spikelets usually more than 1/2 as wide as long; plants green or yellow-green.
→ 2
2. Spikelets 1.9-2.4 mm long; apices of the upper glumes and lower lemmas straight; secondary branches and pedicels divergent; blades often 6-12 mm wide, those of the flag leaves usually more than 1/2 as long as the panicles
subsp. gattingeri
2. Spikelets 1.4-2.1 mm long; apices of the upper glumes and lower lemmas curving over the upper florets at maturity; secondary panicle branches and pedicels appressed; blades usually 2-6 mm wide, those of the flag leaves usually less than M as long as the panicles
subsp. philadelphicum
Source FNA vol. 25, p. 478. FNA vol. 25, p. 459.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Agrostoidea > sect. Agrostoidea Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. amarum, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. anceps subsp. anceps, P. anceps subsp. rhizomatum
P. philadelphicum subsp. gattingeri, P. philadelphicum subsp. lithophilum, P. philadelphicum subsp. philadelphicum
Synonyms P. tuckermanii
Name authority Michx. Bernh. ex Trin.
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