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bitter beachgrass, bitter panicgrass, bitter panicum

panic raide, switch grass, switch panicgrass

Habit Plants perennial; rhizomatous, rhizomes stout, glabrous and glaucous throughout. Plants perennial; rhizomatous, rhizomes often loosely interwoven, hard, with closely overlapping scales, sometimes short or forming a knotty crown.
Culms

20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;

nodes glabrous;

internodes glabrous, glaucous.

40-300 cm tall, 3-5 mm thick, solitary or forming dense clumps, erect or decumbent, usually simple;

nodes glabrous;

internodes hard, glabrous or glaucous, green or purplish.

Sheaths

shorter or longer than the internodes, not keeled, glabrous;

collars often glaucous and purplish;

ligules 1-5 mm;

blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.

longer than the lower internodes, shorter than those above, glabrous or pilose, especially on the throat, margins usually ciliate;

ligules 2-6 mm;

blades 10-60 cm long, 2-15 mm wide, flat, erect, ascending or spreading, glabrous or pubescent, adaxial surfaces sometimes densely pubescent, particularly basally, bases rounded to slightly narrowed, margins scabrous.

Panicles

10-80 cm long, 2-17 cm wide, contracted, slightly nodding;

primary branches whorled or opposite, strongly ascending to appressed;

pedicels 0.5-15 mm, appressed to slightly divergent.

10-55 cm long, 4-20 cm wide, exserted, open;

primary branches thin, straight, solitary to whorled or fascicled, ascending to spreading, scabrous, usually rebranching once;

pedicels 0.5-20 mm, appressed to spreading.

Spikelets

4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;

lower florets staminate.

2.5-8 mm long, 1.2-2.5 mm wide, narrowly lanceoloid, turgid to slightly laterally compressed, glabrous, acuminate.

Glumes

and lower lemmas relatively thick;

lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;

upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;

upper glumes 3.9-7.6 mm, 5-9-veined;

lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;

lower florets staminate;

upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.

Lower

glumes 1.8-3.2 mm, 1/2 - 4/5 as long as the spikelets, glabrous, 5-9-veined, acuminate;

upper glumes and lower lemmas extending 0.4-3 mm beyond the upper florets, 7-11-veined, strongly gaping at the apices;

lower florets staminate;

lower paleas 3-3.5 mm, ovate-hastate, lateral lobes folded over the anthers before anthesis;

upper florets 2.3-3 mm long, 0.8-1.1 mm wide, narrowly ovoid, smooth, glabrous, shiny;

upper lemmas clasping the paleas only at the base.

2n

= 36, 54.

= 18, 21, 25, 30, 32, 35, 36, 54-60, 67-72, 74, 77, 90, 108.

Panicum amarum

Panicum virgatum

Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; MS; NC; NJ; NM; NY; PA; RI; SC; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; WY; HI; MB; NS; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum virgatum grows in tallgrass prairies, especially mesic to wet types where it is a major component of the vegetation, and on dry slopes, sand, open oak or pine woodlands, shores, river banks, and brackish marshes. Its range extends, primarily on the eastern side of the Rocky Mountains, from southern Canada through the United States to Mexico, Cuba, Bermuda, and Costa Rica, and, possibly as an introduction, in Argentina. It has also been introduced as a forage grass to other parts of the world.

Panicum virgatum is an important and palatable forage grass, but its abundance in native grasslands decreases with grazing. Several types are planted for range and wildlife habitat improvement. Plants from eastern New Mexico, western Texas, and northern Mexico tend to have larger spikelets (6-8 mm versus 2.5-5.5 mm) and are sometimes called P. havardii Vasey Tetraploids appear to be the most common ploidy level, especially in the upper midwest and northern plains, with higher ploidy levels being more common southwards, but plants in a small area can range from diploid through duodecaploid, with dysploid derivatives. If morphological markers matched chromosome numbers and ecotypic characters, the species could be considered an aggregate of numerous microspecies. In the absence of such correlations, it must be regarded as simply a wide-ranging, highly variable taxon. Plants identified as Panicum virgatum var. cubense Griseb. and P. virgatum var. spissum Linder represent end points of geographic clines.

Panicum virgatum is not always readily separable from P. amarum, particularly P. amarum subsp. amarulum; future work may support their treatment as conspecific taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long
subsp. amarulum
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long
subsp. amarum
Source FNA vol. 25, p. 472. FNA vol. 25, p. 474.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia
Sibling taxa
P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum
Subordinate taxa
P. amarum subsp. amarulum, P. amarum subsp. amarum
Synonyms P. virgatum var. spissum, P. virgatum var. cubense, P. bavardii
Name authority Elliott L.
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