Panicum amarum
Panicum rigidulum
bitter beachgrass, bitter panicgrass, bitter panicum
redtop panic grass, redtop panicum, smooth witchgrass
20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;
nodes glabrous;
internodes glabrous, glaucous.
35-150 cm, stout, compressed.
shorter or longer than the internodes, not keeled, glabrous;
collars often glaucous and purplish;
ligules 1-5 mm;
blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.
more or less strongly compressed or keeled, sides usually glabrous or sparsely pubescent distally;
ligules 0.3-3 mm, membranous, erose or ciliate, cilia often themselves fimbriate;
blades 8-50 cm long, 2-12 mm wide, flat or folded, both surfaces usually glabrous or scabridulous, or the adaxial surfaces sparsely pilose basally.
10-80 cm long, 2-17 cm wide, contracted, slightly nodding;
primary branches whorled or opposite, strongly ascending to appressed;
pedicels 0.5-15 mm, appressed to slightly divergent.
terminal and axillary, 9-40 cm, 1/3 – 3/4 as wide as long, usually dense; ultimate branchlets usually appressed, 1-sided, scabridulous;
pedicels 0.5-1.5 mm, usually appressed, sometimes with 1-several slender hairs at the apices.
4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;
lower florets staminate.
usually 1.6-3.8 mm, usually subsessile, lanceolate, green, purple-tinged, or purple, glabrous.
and lower lemmas relatively thick;
lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;
upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;
upper glumes 3.9-7.6 mm, 5-9-veined;
lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;
lower florets staminate;
upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.
glumes 2/5 – 3/4 as long as the spikelets, 3-veined, midveins keeled;
upper glumes and lower lemmas subequal or the glumes slightly longer, often spreading slightly apart at the apices, midveins keeled, usually scabridulous apically;
lower florets sterile;
lower paleas to 2/3 as long as the lower lemmas;
upper florets 1.4-2 mm long, 0.6-0.8 mm wide, 2/5 – 3/4 as long as the spikelets, occasionally stipitate, lustrous, with a tuft of minute, thickish hairs at the apices;
upper lemmas thick, stiff, clasping the upper paleas throughout their length.
= 36, 54.
=18.
Panicum amarum
Panicum rigidulum
Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Panicum rigidulum grows in swamps, wet woodlands, flood-plain forests, wet pine savannahs, marshy shores of rivers, ponds, and lakes, drainage ditches, and other similar wet to moist places; it is rarely found in dry sites. Its range extends from southern Canada to Mexico, Guatemala, and the Antilles.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long | subsp. amarulum |
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long | subsp. amarum |
1. Sheaths truncate or broadly auriculate; blade bases much narrower than the subtending sheaths | subsp. abscissum |
1. Sheaths not truncate or broadly auriculate; blade bases about as wide as the subtending sheaths. | → 2 |
2. Blades usually 5-12 mm wide, flat, mostly glabrous or scabridulous; ligules membranous, 0.3-1 mm long. | → 3 |
3. Spikelets 1.6-2.5 mm long, usually over 0.6 mm wide, green or purplish-tinged | subsp. rigidulum |
3. Spikelets 2.4-3 mm long, usually less than 0.6 mm wide, conspicuouly stipitate, usually purple | subsp. elongatum |
2. Blades usually 2-7 mm wide, often folded or involute, usually pilose adaxially, at least near the base; ligules membranous, the cilia usually fimbriate, 0.5-3 mm long. | → 4 |
4. Spikelets 2-2.7 mm long, green or purplish-stained, often obliquely set on the pedicels | subsp. pubescens |
4. Spikelets 2.6-3.8 mm long, usually purple, slender, erect on the pedicels | subsp. combsii |
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