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bitter beachgrass, bitter panicgrass, bitter panicum

Mexican panicgrass, roughstalk witchgrass, roughstalk wltchgrass, woodland panic

Habit Plants perennial; rhizomatous, rhizomes stout, glabrous and glaucous throughout. Plants annual; glabrous or hispid, hairs papillose-based.
Culms

20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;

nodes glabrous;

internodes glabrous, glaucous.

11-110 cm, erect to decumbent;

nodes shortly hirsute or glabrous.

Sheaths

shorter or longer than the internodes, not keeled, glabrous;

collars often glaucous and purplish;

ligules 1-5 mm;

blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.

shorter than the internodes, greenish to purplish, glabrous or with papillose-based hairs, ciliate on 1 margin, glabrous on the other;

collars hirsute;

ligules 1.5-3.5 mm, of hairs;

blades 3-30 cm long, 3-30 mm wide, flat, usually hirsute or sparsely pubescent, hairs papillose-based, sometimes glabrous, bases rounded to cordate-clasping, margins ciliate, cilia papillose-based, apices acute.

Panicles

10-80 cm long, 2-17 cm wide, contracted, slightly nodding;

primary branches whorled or opposite, strongly ascending to appressed;

pedicels 0.5-15 mm, appressed to slightly divergent.

9-30 cm long, 5-8 cm wide, erect or nodding, partially included to well-exserted, rachises glabrous or sparsely hispid basally;

primary branches usually alternate to opposite, divergent, secondary branches and pedicels confined to the distal 2/3;

pulvini inconspicuous;

secondary branches appressed;

pedicels 9-27 mm, appressed.

Spikelets

4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;

lower florets staminate.

1.9-4 mm long, 0.8-1 mm wide, ovoid to almost spherical, often reddish-brown, glabrous, veins prominent, scabridulous, apices abruptly acuminate.

Glumes

and lower lemmas relatively thick;

lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;

upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;

upper glumes 3.9-7.6 mm, 5-9-veined;

lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;

lower florets staminate;

upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.

Lower

glumes 1.3-2.4 mm, 1/2 - 3/4 as long as the spikelets, 3-5-veined;

upper glumes 1.8-3.3 mm, 7-11-veined;

lower florets sterile;

lower lemmas similar to the upper glumes, 9-veined;

lower paleas 0.4-0.9 mm;

upper florets 1.5-2.4 mm long, 0.4-0.8 mm wide, ellipsoid, smooth or conspicuously papillate, shiny, stramineous, often with a lunate scar at the base.

2n

= 36, 54.

Panicum amarum

Panicum hirticaule

Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; MS; NC; NJ; NM; NY; PA; RI; SC; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; OK; TX; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum hirticaule grows in rocky or sandy soils in waste places, roadsides, ravines, and wet meadows along streams. Its range extends from southeastern California and southwestern Texas southward through Mexico, Central America, Cuba, and Hispaniola to western South America and Argentina.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long
subsp. amarulum
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long
subsp. amarum
1. Blades rounded at the base, 3-16 mm wide; lower paleas less than 1/2 as long as the upper florets; panicles erect
subsp. hirticaule
1. Blades cordate, clasping at the base, 4-30 mm wide; lower paleas more than 1/2 as long as the upper florets; panicles often nodding.
→ 2
2. Nodes, sheaths, and blades glabrous or sparsely pilose, hairs papillose-based; culms usually less than 70 cm tall; spikelets 3.2-4 mm long
subsp. stramineum
2. Nodes, sheaths, and blades hirsute, hairs papillose-based; culms robust, usually more than 70 cm tall; spikelets 3-3.3 mm long
subsp. sonorum
Source FNA vol. 25, p. 472. FNA vol. 25, p. 460.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. amarum subsp. amarulum, P. amarum subsp. amarum
P. hirticaule subsp. hirticaule, P. hirticaule subsp. sonorum, P. hirticaule subsp. stramineum
Synonyms P. pampinosum
Name authority Elliott J. Presl
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