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bitter beachgrass, bitter panicgrass, bitter panicum

panic flexible, wiry panic grass, wiry witch grass

Habit Plants perennial; rhizomatous, rhizomes stout, glabrous and glaucous throughout. Plants annual; delicate, green or yellow-green.
Culms

20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;

nodes glabrous;

internodes glabrous, glaucous.

10-75 cm, about 1 mm thick, simple or with erect basal branches;

nodes densely pilose, hairs ascending;

internodes glabrous or shortly pubescent distally.

Sheaths

shorter or longer than the internodes, not keeled, glabrous;

collars often glaucous and purplish;

ligules 1-5 mm;

blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.

longer than the internodes, green to purplish, hispid, margins sparsely ciliate;

ligules 0.5-1.5 mm;

blades 3-32 cm long, 1-7 mm wide, ascending to erect, linear, narrowing basally, flat or the margins involute, surfaces sparsely hirsute or pilose (rarely glabrous), hairs near the base papillose-based, margins prominent, apices acute.

Panicles

10-80 cm long, 2-17 cm wide, contracted, slightly nodding;

primary branches whorled or opposite, strongly ascending to appressed;

pedicels 0.5-15 mm, appressed to slightly divergent.

5-45 cm long, 1-6 cm wide, at least 1/2 as long as the plants and 3 times longer than wide, open;

rachises glabrous;

primary branches usually alternate or subopposite, ascending to slightly divergent, secondary branches and pedicels attached to the distal 2/3;

secondary branches diverging;

pedicels 0.5-17 mm, ascending to appressed.

Spikelets

4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;

lower florets staminate.

2.5 - 3.7 mm long, 0.6-1.1 mm wide, narrowly ovoid, glabrous, acute;

lower glumes 0.8-1.3 mm, 1/3 – 1/2 as long as the spikelets, acuminate;

upper glumes 2.3-3.3 mm, 7-9-veined, exceeding the upper florets by about 0.6 mm;

lower florets sterile;

lower lemmas 2.2-2.7 mm, exceeding the upper florets by about 0.6 mm, 7- or 9-veined, apices scabridulous, pointed;

lower paleas absent;

upper florets 1.6-1.7 mm long, about 0.6 mm wide, usually smooth, usually pale, sometimes becoming dark at maturity.

Glumes

and lower lemmas relatively thick;

lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;

upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;

upper glumes 3.9-7.6 mm, 5-9-veined;

lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;

lower florets staminate;

upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.

2n

= 36, 54.

= 18.

Panicum amarum

Panicum flexile

Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; MS; NC; NJ; NM; NY; PA; RI; SC; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NJ; NY; OH; OK; PA; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum flexile grows in fens and other calcareous wetlands, in dry, calcareous or mafic rock barrens, and in open woodlands, especially on limestone derived soils. It is restricted to the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long
subsp. amarulum
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long
subsp. amarum
Source FNA vol. 25, p. 472. FNA vol. 25, p. 460.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. amarum subsp. amarulum, P. amarum subsp. amarum
Name authority Elliott (Gatt.) Scribn.
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