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bitter beachgrass, bitter panicgrass, bitter panicum

fall panicgrass, fall panicum, knee grass, panic d'automne, smooth witchgrass

Habit Plants perennial; rhizomatous, rhizomes stout, glabrous and glaucous throughout. Plants annual or short-lived perennials in the Flora region, perennial in the tropics; usually terrestrial, sometimes aquatic but not floating.
Culms

20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;

nodes glabrous;

internodes glabrous, glaucous.

5-200 cm tall, 0.4-3 mm thick, decumbent to erect, commonly geniculate to ascending, rooting at the lower nodes when in water, simple to divergently branched from the lower and middle nodes, usually succulent, slightly compressed, glabrous;

nodes usually swollen, sometimes constricted on robust plants, glabrous;

internodes glabrous, shiny, pale green to purplish.

Sheaths

shorter or longer than the internodes, not keeled, glabrous;

collars often glaucous and purplish;

ligules 1-5 mm;

blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.

compressed, inflated, sparsely pubescent near the base, elsewhere mostly glabrous, sparsely pilose, or hispid, hairs sometimes papillose-based, margins or throat ciliate, with papillose-based hairs;

ligules 0.5-2 mm;

blades 10-65 cm long, 3-25 mm wide, glabrous or sparsely pilose, often scabrous near the margins, midribs stout, whitish.

Panicles

10-80 cm long, 2-17 cm wide, contracted, slightly nodding;

primary branches whorled or opposite, strongly ascending to appressed;

pedicels 0.5-15 mm, appressed to slightly divergent.

4-40 cm, diffuse, lax, with a few spikelets;

branches to 15 cm, alternate or opposite, occasionally verticillate, ascending to spreading, stiff, scabrous;

pedicels 1-6 mm, sharply 3-angled, scabrous, expanded to cuplike apices, appressed mostly to the abaxial side of the branches.

Spikelets

4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;

lower florets staminate.

1.8-3.8 mm long, 0.7-1.2 mm wide, ellipsoid to narrowly ovoid, light green to red-purple, glabrous, acute to acuminate.

Glumes

and lower lemmas relatively thick;

lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;

upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;

upper glumes 3.9-7.6 mm, 5-9-veined;

lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;

lower florets staminate;

upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.

Lower

glumes 0.6-1.2 mm, 1/4 - 1/3 as long as the spikelets, 0-3-veined, obtuse to acute;

upper glumes and lower lemmas similar, exceeding the upper florets by 0.3-0.6 mm, 7-9-veined;

lower paleas vestigial to almost as long as the lower lemmas;

lower florets sterile;

upper florets 1.4-2.5 mm long, 0.7-1.1 mm wide, narrowly ellipsoid, smooth, shiny, stramineous to nigrescent, with pale veins.

2n

= 36, 54.

= 36, 54.

Panicum amarum

Panicum dichotomiflorum

Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; MS; NC; NJ; NM; NY; PA; RI; SC; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; HI; PR; BC; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum dichotomiflorum grows in open, often wet, disturbed areas such as cultivated and fallow fields, roadsides, ditches, open stream banks, receding shores, clearings in flood plain woods, and sometimes in shallow water. It is probably native throughout the eastern United States and adjacent Canada, but introduced elsewhere, including in the western United States. Its size and habit may be partly under genetic control, but these features also seem to be strongly affected by moisture levels, soil richness, competition, and the time of germination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long
subsp. amarulum
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long
subsp. amarum
1. Spikelets 1.8-2.2 mm long, widest at the middle, acute; upper glumes and lower lemmas submembranaceous; pedicels often over 3 mm long
subsp. puritanorum
1. Spikelets 2.2-3.8 mm long, widest below the middle, acuminate; upper glumes and lower lemmas subcoriaceous; most pedicels less than 3 mm long.
→ 2
2. Sheaths glabrous or sparsely pilose, hairs not papillose-based
subsp. dichotomiflorum
2. Sheaths hispid, hairs papillose-based
subsp. bartowense
Source FNA vol. 25, p. 472. FNA vol. 25, p. 469.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Dichotomiflora
Sibling taxa
P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. amarum subsp. amarulum, P. amarum subsp. amarum
P. dichotomiflorum subsp. bartowense, P. dichotomiflorum subsp. dichotomiflorum, P. dichotomiflorum subsp. puritanorum
Name authority Elliott Michx.
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