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bitter beachgrass, bitter panicgrass, bitter panicum

common panicgrass, common witchgrass, old witch grass, panic capillaire, witch grass, witch panicgrass

Habit Plants perennial; rhizomatous, rhizomes stout, glabrous and glaucous throughout. Plants annual; hirsute or hispid, hairs papillose-based, often bluish or purplish.
Culms

20-250 cm tall, 3-10 mm thick, erect or decumbent, simple or branched from the lower nodes;

nodes glabrous;

internodes glabrous, glaucous.

15-130 cm, slender to stout, not woody, erect to decumbent, straight to zigzag, simple to profusely branched;

nodes sparsely to densely pilose.

Sheaths

shorter or longer than the internodes, not keeled, glabrous;

collars often glaucous and purplish;

ligules 1-5 mm;

blades 7-50 cm long, 2-13 mm wide, erect or ascending, firm, thick, flat basally, more or less involute towards the apices.

rounded, hirsute or hispid, hairs papillose-based;

ligules membranous, ciliate, cilia 0.5-1.5 mm;

blades 5-40 cm long, 3-18 mm wide, linear, spreading.

Panicles

10-80 cm long, 2-17 cm wide, contracted, slightly nodding;

primary branches whorled or opposite, strongly ascending to appressed;

pedicels 0.5-15 mm, appressed to slightly divergent.

13-50 cm long, 7-24 cm wide, usually more than 1/2 as long as the plants, included at the base or exserted at maturity, disarticulating at the base of the peduncles at maturity and becoming a tumbleweed;

branches spreading;

pedicels 0.5-2.8 mm, scabrous, pilose.

Spikelets

4-7.7 mm long, 1.5-2 mm wide, narrowly ovoid, glabrous, acuminate;

lower florets staminate.

1.9-4 mm, ellipsoid to lanceoloid, often red-purple, glabrous.

Glumes

and lower lemmas relatively thick;

lower glumes 2.8-4 mm, 1/2 - 4/5 as long as the spikelets, 3-9-veined, apices of the midveins sometimes scabridulous;

upper glumes and lower lemmas extending 1.5-3 mm beyond the upper florets, apices stiffly gaping;

upper glumes 3.9-7.6 mm, 5-9-veined;

lower lemmas slightly shorter than the upper glumes, 7-9-veined, lower paleas 3-7 mm, oblong-hastate, folded over the anthers;

lower florets staminate;

upper florets 2.4-3.9 mm long, 1-1.8 mm wide, narrowly ovoid to oblong, glabrous, smooth, shiny, lemma margins clasping the paleas only at the base.

Lower

florets sterile;

lower glumes 1/3– 1/2 as long as the spikelets, 1-3-veined;

upper glumes 1.8-3.1 mm, 7-9-veined, midveins scabridulous;

lower lemmas 1.9-3 mm, extending 0.4-1.1 mm beyond the upper florets, often stiff, straight, prominently veined distally;

upper florets stramineous or nigrescent, sometimes with a prominent lunate scar at the base, often disarticulating before the glumes, leaving the empty glumes and lower lemmas temporarily persisting on the panicles.

2n

= 36, 54.

= 18.

Panicum amarum

Panicum capillare

Distribution
from FNA
AL; CT; DE; FL; GA; LA; MA; MD; MS; NC; NJ; NM; NY; PA; RI; SC; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Virgin Islands
[WildflowerSearch map]
[BONAP county map]
Discussion

Panicum amarum grows in the coastal dunes, wet sandy soils, and the margins of swamps, along the Atlantic Ocean and the Gulf of Mexico from Connecticut to northeastern Mexico. It is also known, as an introduction, from a few inland locations in New Mexico, North Carolina, and West Virginia, as well as in the Bahamas and Cuba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Panicum capillare grows in open areas, particularly in disturbed sites such as fields, pastures, roadsides, waste places, ditches, sand, and rock crevices, etc. It grows throughout temperate North America, including northern Mexico. It also grows in Bermuda, the Virgin Islands, and sporadically in South America, and has become naturalized in much of Europe and Asia. It appears to hybridize with P. philadelphicum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Rhizomes short or ascending; culms often bunched and decumbent, usually more than 120 cm tall; lower glumes with 3-5 less evident veins, the midvein smooth distally; spikelet density high; panicles with 2 or more main branches per node; spikelets 4-5.9 mm long
subsp. amarulum
1. Rhizomes horizontally elongate; culms mostly solitary, less than 150 cm tall; lower glumes with 7-9 prominent veins, the midvein scabridulous distally; spikelet density moderate; panicles with 1 or 2 main branches per node; spikelets 4.7-7.7 mm long
subsp. amarum
1. Upper florets without a lunate scar, usually stramineous; lower paleas absent; pedicels and secondary branches strongly divergent
subsp. capillare
1. Upper florets with a lunate scar at the base, usually nigrescent; lower paleas present; pedicels and secondary branches often appressed, varying to narrowly divergent
subsp. hillmanii
Source FNA vol. 25, p. 472. FNA vol. 25.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Repentia Poaceae > subfam. Panicoideae > tribe Paniceae > Panicum > subg. Panicum > sect. Panicum
Sibling taxa
P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillare, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
P. amarum, P. anceps, P. antidotale, P. bergii, P. bisulcatum, P. brachyanthum, P. bulbosum, P. capillarioides, P. coloratum, P. dichotomiflorum, P. diffusum, P. flexile, P. ghiesbreghtii, P. gymnocarpon, P. hallii, P. hemitomon, P. hirsutum, P. hirticaule, P. lacustre, P. miliaceum, P. mohavense, P. obtusum, P. paludosum, P. philadelphicum, P. plenum, P. psilopodium, P. repens, P. rigidulum, P. tenerum, P. trichoides, P. urvilleanum, P. verrucosum, P. virgatum
Subordinate taxa
P. amarum subsp. amarulum, P. amarum subsp. amarum
P. capillare subsp. capillare, P. capillare subsp. hillmanii
Name authority Elliott L.
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