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alpine rock butterweed, hoary groundsel

Bolander's groundsel, Bolander's ragwort

Habit Perennials, 7–15+ cm; ± rhizomatous (rhizomes branched, sometimes densely crowded). Perennials, 20–50+ cm; fibrous-rooted (caudices creeping).
Stems

1 or 3–5, clustered (often scapiform), usually floccose, lanate-tomentose, or canescent, sometimes glabrate.

1 or 2–3, loosely clustered, glabrous.

Basal leaves

(either of two forms): usually (1) sessile, sometimes petiolate;

blades narrowly lanceolate to elliptic, 15–40+ × 5–25 mm, bases tapering, margins entire or dentate toward apices (often revolute), sometimes (2) petiolate;

blades ovate to orbiculate, 10–20 × 5–15 mm, bases tapering to abruptly contracted, margins entire or wavy, sometimes dentate toward apices.

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

Cauline leaves

abruptly reduced (bractlike).

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

Peduncles

inconspicuously bracteate, glabrous or densely hairy.

ebracteate or inconspicuously bracteate, glabrous.

Ray florets

0, 8, or 13;

corolla laminae 5–10 mm.

8 or 13;

corolla laminae 10–15+ mm.

Disc florets

30–50+;

corolla tubes 2.5–3.5 mm, limbs 3–4 mm.

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

Phyllaries

13 or 21, green (tips sometimes cyanic), 4–10 mm, glabrous or hairy.

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

Calyculi

conspicuous (bractlets often cyanic).

conspicuous.

Heads

1–5(–8) in cymiform to subumbelliform arrays.

8–15+ in corymbiform or subumbelliform arrays.

Cypselae

1.5–2 mm, glabrous;

pappi 5–6 mm.

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

2n

= 44, 46.

= 46.

Packera werneriifolia

Packera bolanderi

Phenology Flowering mid Jun–mid Aug.
Habitat Rocky talus slopes, sandy soils in forest openings near or above timberline
Elevation 2400–3700 m (7900–12100 ft)
Distribution
from FNA
AZ; CA; CO; ID; NM; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Packera werneriifolia is morphologically variable; it occurs throughout the central Rockies and, sporadically, as far west as the Sierra Nevada. Leaf morphology varies from ovate, elliptic, or narrowly elliptic in the Rockies to narrow with revolute margins in California and Arizona. All specimens are characteristically scapiform.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 602. FNA vol. 20, p. 579.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio aureus var. werneriifolius, Senecio alpicola, Senecio molinarius, Senecio muirii, Senecio perennans, Senecio petraeus, Senecio petrocallis, Senecio petrophilus, Senecio saxosus, Senecio scaposus, Senecio werneriifolius Senecio bolanderi
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 48. (1981) (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981)
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