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threetooth ragwort

Bolander's groundsel, Bolander's ragwort

Habit Perennials, 10–30+ cm; taprooted (caudices ascending to erect). Perennials, 20–50+ cm; fibrous-rooted (caudices creeping).
Stems

1 or multiple, clustered, usually glabrous, rarely sparsely floccose-tomentose, leaf axils sometimes tomentose.

1 or 2–3, loosely clustered, glabrous.

Basal leaves

(and proximal cauline, relatively thick and turgid) petiolate;

blades lanceolate or narrowly oblanceolate, 20–40+ × 5–15 mm, bases tapering, margins usually entire, sometimes sub-pinnatisect (apices subentire or dentate).

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

Cauline leaves

gradually reduced (± petiolate or sessile).

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

Peduncles

bracteate, glabrous or sparsely tomentose.

ebracteate or inconspicuously bracteate, glabrous.

Ray florets

8–10(–13);

corolla laminae 5–8+ mm.

8 or 13;

corolla laminae 10–15+ mm.

Disc florets

45–60+;

corolla tubes 3–3.5 mm, limbs 4–5 mm.

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

Phyllaries

13 or 21, green, 6–10 mm, sparsely tomentose proximally, glabrous distally.

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

Calyculi

inconspicuous.

conspicuous.

Heads

4–15+ in corymbiform arrays.

8–15+ in corymbiform or subumbelliform arrays.

Cypselae

1.5–2.5 mm, glabrous or sparsely hirtellous on ribs;

pappi 5–6 mm.

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

2n

= 46.

= 46.

Packera tridenticulata

Packera bolanderi

Phenology Flowering late May–early Jul.
Habitat Open, dry areas, roadsides, gravelly or sandy slopes, short-grass prairies or sagebrush scrubs
Elevation 1000–2000 m (3300–6600 ft)
Distribution
from FNA
CO; KS; ND; NE; NM; OK; SD; TX; WY
[WildflowerSearch map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Packera tridenticulata is found throughout the central High Plains, in the foothills of the Rocky Mountains, and in high valleys to the west of the Rocky Mountain front. It tends to grow in clumps; the multiple stems arise from well-developed taproots. It apparently hybridizes with P. neomexicana var. mutabilis where their ranges overlap.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 601. FNA vol. 20, p. 579.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio tridenticulatus, Senecio acutidens, Senecio compactus, Senecio densus, Senecio oblanceolatus Senecio bolanderi
Name authority (Rydberg) W. A. Weber & Á. Löve: Phytologia 49: 48. (1981) (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981)
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