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balsam groundsel, Canadian butterweed

splitleaf groundsel

Habit Perennials, 20–45+ cm; subrhizomatous (bases weakly branched, ascending to erect). Perennials, 10–30+ cm; fibrous-rooted (caudices horizontal to erect, relatively short and stout).
Stems

1 or 2–4, loosely clustered, glabrous or sparsely tomentose proximally.

usually 1, rarely 2–3, clustered, glabrous.

Basal leaves

petiolate;

blades lanceolate to narrowly elliptic or oblanceolate, 30–60+ × 10–20+ mm, bases tapering, sometimes obtuse, margins subentire to dentate or serrate.

(relatively thick and turgid) petiolate (petioles broad-winged or narrow, delicate);

blades ovate, ovate-orbiculate, oblong-lanceolate, orbiculate, suborbiculate, reniform, or subreniform, 10–50+ × 10–40 mm, bases tapering or subcordate to abruptly contracted, margins entire or crenate.

Cauline leaves

gradually reduced (proximals petiolate, sublyrate; mids sessile, not clasping, lanceolate, dissected, incised, or lacerate; distals sessile, bractlike).

gradually reduced (proximal and mid sometimes as large as or larger than basal, sessile, oblanceolate to obovate, sometimes lyrate, sometimes auriculate and/or clasping, subentire to irregularly dissected; distal usually sessile, bractlike, sometimes larger).

Peduncles

usually bracteate, sometimes ebracteate, glabrous.

inconspicuously bracteate or ebracteate, glabrous.

Ray florets

0, 8, or 13;

corolla laminae (pale yellow) 5–10+ mm.

8 or 13;

corolla laminae (deep yellow to orange) 5–8+ mm.

Disc florets

50–65+;

corolla tubes 2–3 mm, limbs 2–3 mm.

45–60+;

corolla tubes 2–3 mm, limbs 2.5–3.5 mm.

Phyllaries

13 or 21, green, 5–8+ mm, glabrous.

(8–)13 or 21, green (tips cyanic), 5–7+ mm, glabrous.

Calyculi

inconspicuous.

conspicuous (bases of bractlets swollen).

Heads

2–10+ in loose or compact, corymbiform arrays.

1–6+ in congested, corymbiform or subumbelliform arrays.

Cypselae

1–2 mm, usually glabrous, sometimes hispidulous on ribs;

pappi 3.5–4.5 mm.

0.75–1.5 mm, glabrous, pappi 3–4 mm.

2n

= 44, 46, 92.

= 46.

Packera paupercula

Packera dimorphophylla

Phenology Flowering May–late Jun (south), late Jun–early Aug (north).
Habitat Wet meadows, open woodlands, along streams, rocky outcrops
Elevation 0–3000 m (0–9800 ft)
Distribution
from FNA
AK; AL; CO; DC; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; PA; SD; TN; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; NM; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Ecologically and morphologically, Packera paupercula is the most variable species of the genus in North America. Some “phases” have been treated as separate species, subspecies, varieties, forms, and races. Variation within P. paupercula hints at some interesting evolutionary relationships; characteristics used to separate taxa overlap. Much of the morphologic variation in this species may be due to hybridization and introgression. I do not recognize any of the infraspecific taxa that have been proposed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves: petioles broad-winged, blades ovate to oblong-lanceolate, bases tapering; stems usually not branched
var. dimorphophylla
1. Basal leaves: petioles narrowly, if at all winged, blades suborbiculate, orbiculate, ovate-orbiculate, subreniform, or reniform, bases subcordate to abruptly contracted; stems branched or not branched
→ 2
2. Plants 30–50 cm; stems not branched; petiole lengths 1–3 times blades; Utah
var. intermedia
2. Plants 10–35 cm; stems frequently 1–3 times branched; petiole lengths 1–2 times blades; Idaho, Montana, Nevada, Wyoming
var. paysonii
Source FNA vol. 20, p. 595. FNA vol. 20, p. 584.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. dimorphophylla var. dimorphophylla, P. dimorphophylla var. intermedia, P. dimorphophylla var. paysonii
Synonyms Senecio pauperculus, Senecio balsamitae, Senecio balsamitae var. firmifolius, Senecio balsamitae var. thomsoniensis, Senecio crawfordii, Senecio flavovirens, Senecio gaspensis, Senecio multnomensis, Senecio robbinsii var. subtomentosus, Senecio tweedyi Senecio dimorphophyllus
Name authority (Michaux) Á. Löve & D. Löve: Bot. Not. 128: 520. (1976) (Greene) W. A. Weber & Á. Löve: Phytologia 49: 46. (1981)
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