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Bolander's groundsel, Bolander's ragwort

Oak Creek ragwort

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials, 60–100+ cm; taprooted (caudices subligneous, ascending to erect).
Stems

1 or 2–3, loosely clustered, glabrous.

1 or 2–4, clustered (proximally deeply purple-tinged, distally lightly tinged), glabrous or tomentose at bases and in leaf axils.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(and proximal cauline) petiolate;

blades obovate or lyrate (pinnately lobed, lateral lobes 2–6+ pairs, their bases petioluliform, terminal lobes larger than laterals, midribs narrowly winged), 60–160+ × 20–40+ mm, bases wide, ultimate margins sharply dentate, crenate-dentate, or irregularly incised.

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually reduced (petiolate or sessile; shallowly lobed, midribs ± winged, distals bractlike, dentate to incised).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

ebracteate, glabrous.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

(8–)13;

corolla laminae 6–10+ mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

60–70+;

corolla tubes 2–3 mm, limbs 3.5–4.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

(13–)21, green (tips yellow), 5–7 mm, glabrous (tips sometimes hairy).

Calyculi

conspicuous.

inconspicuous.

Heads

8–15+ in corymbiform or subumbelliform arrays.

15–40+ in open, cymiform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

1.5–2 mm, glabrous or ± scabrellous;

pappi 5.5–6.5 mm.

2n

= 46.

= 92.

Packera bolanderi

Packera quercetorum

Phenology Flowering mid Apr–early Jun.
Habitat Rocky soils, open areas, scrub-oak and pinyon-pine forests, chaparral
Elevation 800–2200 m (2600–7200 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera quercetorum is found only infrequently and in relatively small populations in central and southern Arizona and west-central New Mexico. The plants are robust and have probable affinities to P. multilobata. The plants have a bluish tinge when freshly collected and are distinctive in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 598.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio quercetorum, Senecio macropus
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Greene) C. Jeffrey: Kew Bull. 47: 101. (1992)
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